Estuarine Fish
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Coastal fish, also called inshore fish or neritic fish, inhabit the sea between the
shoreline A coast (coastline, shoreline, seashore) is the land next to the sea or the line that forms the boundary between the land and the ocean or a lake. Coasts are influenced by the topography of the surrounding landscape and by aquatic erosion, su ...
and the edge of the
continental shelf A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an islan ...
. Since the continental shelf is usually less than deep, it follows that
pelagic The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the sur ...
coastal fish are generally
epipelagic fish Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that live on or near the bottom, and reef fish that are associated with coral reefs. ...
, inhabiting the sunlit
epipelagic zone The photic zone (or euphotic zone, epipelagic zone, or sunlight zone) is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological ...
.Moyle and Cech, 2004, page 585 Coastal fish can be contrasted with ''oceanic fish'' or ''offshore fish'', which inhabit the deep seas beyond the continental shelves. Coastal fish are the most abundant in the world. They can be found in
tidal pool A tide pool or rock pool is a shallow pool of seawater that forms on the rocky intertidal shore. These pools typically range from a few inches to a few feet deep and a few feet across. Many of these pools exist as separate bodies of water only ...
s,
fjord In physical geography, a fjord (also spelled fiord in New Zealand English; ) is a long, narrow sea inlet with steep sides or cliffs, created by a glacier. Fjords exist on the coasts of Antarctica, the Arctic, and surrounding landmasses of the n ...
s and
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
, near sandy shores and rocky coastlines, around
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in group ...
s and on or above the continental shelf. Coastal fish include
forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish that feed on planktons (i.e. planktivores) and other small aquatic organisms (e.g. krill). They are in turn preyed upon by various predators including larger fish, seabirds ...
and the
predator fish Predatory fish are hypercarnivorous fish that actively prey upon other fish or aquatic animals, with examples including shark, billfish, barracuda, alligator gar, tuna, dolphinfish, walleye, perch and salmon. Some omnivorous fish, such as t ...
that feed on them. Forage fish thrive in inshore waters where high
productivity Productivity is the efficiency of production of goods or services expressed by some measure. Measurements of productivity are often expressed as a ratio of an aggregate output to a single input or an aggregate input used in a production proce ...
results from
upwelling Upwelling is an physical oceanography, oceanographic phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water from deep water towards the ocean surface. It replaces the warmer and usually nutrient-depleted sur ...
and shoreline run off of nutrients. Some are partial residents that spawn in streams, estuaries and bays, but most complete their life cycles in the zone.Moyle and Cech, 2004, page 572


Coastal habitats

Coastal fish are found in the waters above the
continental shelves A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...
that extend from the continental
shoreline A coast (coastline, shoreline, seashore) is the land next to the sea or the line that forms the boundary between the land and the ocean or a lake. Coasts are influenced by the topography of the surrounding landscape and by aquatic erosion, su ...
s, and around the
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in group ...
s that surround volcanic islands. The total world shoreline extends for World
The World Factbook, CIA. Retrieved 26 February 2010.
and the continental shelves occupy a total area of 24.3 million km2 (9 376 million sq mi). This is nearly 5% of the world's total area of 510 million km2.


Nearshore fish

Nearshore fish, sometimes called littoral fish, live close to the shore. They are associated with the
intertidal zone The intertidal zone or foreshore is the area above water level at low tide and underwater at high tide; in other words, it is the part of the littoral zone within the tidal range. This area can include several types of habitats with various ...
, or with
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
,
lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into ''coastal lagoons'' (or ''barrier lagoons'') an ...
s,
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in group ...
s,
kelp forest Kelp forests are underwater areas with a high density of kelp, which covers a large part of the world's coastlines. Smaller areas of anchored kelp are called kelp beds. They are recognized as one of the most productive and dynamic ecosystems on E ...
s,
seagrass meadow A seagrass meadow or seagrass bed is an underwater ecosystem formed by seagrasses. Seagrasses are marine (saltwater) plants found in shallow coastal waters and in the brackish waters of estuaries. Seagrasses are flowering plants with stems and ...
s, or rocky or sandy bottoms, usually in shallow waters less than about deep.


Intertidal fish

Intertidal fish are fish that move in and out with the
tide Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another. Tide tables ...
in the
intertidal zone The intertidal zone or foreshore is the area above water level at low tide and underwater at high tide; in other words, it is the part of the littoral zone within the tidal range. This area can include several types of habitats with various ...
of the seashore, or are found in
rock pool A tide pool or rock pool is a shallow pool of seawater that forms on the rocky intertidal shore. These pools typically range from a few inches to a few feet deep and a few feet across. Many of these pools exist as separate bodies of water only ...
s or under rocks. The
intertidal zone The intertidal zone or foreshore is the area above water level at low tide and underwater at high tide; in other words, it is the part of the littoral zone within the tidal range. This area can include several types of habitats with various ...
of rocky shores can contain indentations which trap pools of salty water, called
rock pool A tide pool or rock pool is a shallow pool of seawater that forms on the rocky intertidal shore. These pools typically range from a few inches to a few feet deep and a few feet across. Many of these pools exist as separate bodies of water only ...
s. Living in these habitats are communities of hardy plant and animal species specially adapted for coping with the volatile environment around them. The plants and animals interact with each other and with the rock pool to form miniature
ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...
s, easily accessible to students and a source of fascination for young children. Plants such as
seaweed Seaweed, or macroalgae, refers to thousands of species of macroscopic, multicellular, marine algae. The term includes some types of ''Rhodophyta'' (red), '' Phaeophyta'' (brown) and ''Chlorophyta'' (green) macroalgae. Seaweed species such as ...
s, cnidarians such as
sea anemone Sea anemones ( ) are a group of predation, predatory marine invertebrates constituting the order (biology), order Actiniaria. Because of their colourful appearance, they are named after the ''Anemone'', a terrestrial flowering plant. Sea anemone ...
s, arthropods like
barnacle Barnacles are arthropods of the subclass (taxonomy), subclass Cirripedia in the subphylum Crustacean, Crustacea. They are related to crabs and lobsters, with similar Nauplius (larva), nauplius larvae. Barnacles are exclusively marine invertebra ...
s, and
mollusc Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
s such as the common
limpet Limpets are a group of aquatic snails with a conical gastropod shell, shell shape (patelliform) and a strong, muscular foot. This general category of conical shell is known as "patelliform" (dish-shaped). Existing within the class Gastropoda, ...
and the
common periwinkle The common periwinkle or winkle (''Littorina littorea'') is a species of small edible whelk or sea snail, a marine gastropod mollusc that has gills and an operculum, and is classified within the family Littorinidae, the periwinkles. This is a ...
can be permanent residents of rock pools. But most rock pool animals, such as
crab Crabs are decapod crustaceans of the infraorder Brachyura (meaning "short tailed" in Greek language, Greek), which typically have a very short projecting tail-like abdomen#Arthropoda, abdomen, usually hidden entirely under the Thorax (arthropo ...
s,
shrimp A shrimp (: shrimp (American English, US) or shrimps (British English, UK)) is a crustacean with an elongated body and a primarily Aquatic locomotion, swimming mode of locomotion – typically Decapods belonging to the Caridea or Dendrobranchi ...
and fish are just temporary residents, occupying a rock pool only until the next tide takes them to a new location. Some rock pool fish which are temporary residents include the
long-spined sea scorpion The long-spined bullhead (''Taurulus bubalis''), also known as the long-spined sea-scorpion, and the long-spined scorpion fish is a coastal fish of the sculpin family Cottidae, inhabiting marine waters of Europe. Taxonomy The longspined bullhea ...
, the pipefish worm, the rock goby and the common
lumpsucker The Cyclopteridae are a family of marine fishes, commonly known as lumpsuckers or lumpfish, in the order Scorpaeniformes. They are found in the cold waters of the Arctic Ocean, Arctic, Atlantic Ocean, North Atlantic, and Pacific Ocean, North Paci ...
. However some other rock pool fish are territorial in nature, and will stay with the same pool for extended periods. Examples are the common blenny and its near relative the butterfish. * The common
blenny Blennies (from the Greek and , mucus, slime) are a diverse clade of ray-finned fish in the suborder Blennioidei of the percomorph order Blenniiformes. They inhabit marine, brackish, and occasionally freshwater habitats, and generally share sim ...
, also known as the shanny, is found in northern temperate waters. They hide under rocks and in crannies in rock pools when the tide is out. They feed on green seaweed and invertebrates such as barnacles. They can crawl on dry land, using their paired fins. About long, they have smooth skin, without scales, and are covered with soft slime. The slime prevents them drying if they are stranded on a shore between tides. So long as their skin stays moist, they can breathe out of water. They are sometimes called "sea frogs" because they bask in the sun on weeds outside the water, and like frogs, jump to safety when disturbed. They can change their colour to match their surroundings. The female lays eggs in crevices or under stones and the male guards them until they hatch. In the winter, when storms can be severe, they move out of their rock pools into the shallows. The common blenny is bold with strong teeth, and will bite humans if it feels threatened. * The rock goby is a small fish, about , found in northern temperate waters. It is coloured black with white blotches, and hides under stones and amongst seaweed. It is a temporary resident of rock pools when the tide is out. The female rock goby lays eggs on the underside of rocks and shells and then leaves them. The male guards the eggs until they hatch. First-year rock gobies often visit rock pools in winter when the older fish have left. File:Australian blenny.jpg, Australian
blenny Blennies (from the Greek and , mucus, slime) are a diverse clade of ray-finned fish in the suborder Blennioidei of the percomorph order Blenniiformes. They inhabit marine, brackish, and occasionally freshwater habitats, and generally share sim ...
File:Gobius_paganellus.jpg, Rock goby File:Long_Spined_Scorpion_Fish.jpg,
Long-spined sea scorpion The long-spined bullhead (''Taurulus bubalis''), also known as the long-spined sea-scorpion, and the long-spined scorpion fish is a coastal fish of the sculpin family Cottidae, inhabiting marine waters of Europe. Taxonomy The longspined bullhea ...
File:smooth lumpsucker.jpg, Smooth
lumpsucker The Cyclopteridae are a family of marine fishes, commonly known as lumpsuckers or lumpfish, in the order Scorpaeniformes. They are found in the cold waters of the Arctic Ocean, Arctic, Atlantic Ocean, North Atlantic, and Pacific Ocean, North Paci ...
inflated in a defensive response
* The
long-spined sea scorpion The long-spined bullhead (''Taurulus bubalis''), also known as the long-spined sea-scorpion, and the long-spined scorpion fish is a coastal fish of the sculpin family Cottidae, inhabiting marine waters of Europe. Taxonomy The longspined bullhea ...
, a small stout fish which grows about long, is another temporary resident of rocky pools. They have large black eyes, a large mouth, and four long spines—two on each side on the gill cover—that stick out when the fish is removed from the water. They also have an organ like a finger on each side of their mouths which helps them catch prey. Because of their broad heads, they are also called "bullheads". They have a variety of effective camouflaged colours ranging from shades of browns with cream blotches, to orange and red with white blotches. They can also change their body colour to match their surroundings. They are found around the coasts of Northern Europe in shallow rocky waters hiding amongst seaweed. They are also found in rock pools and sometimes in waters deep. Long-spined sea scorpions lay eggs amongst seaweed or attached to rock crevices. The young hatch after two or three weeks, and go through several development stages before maturing into adults. *
Lumpsucker The Cyclopteridae are a family of marine fishes, commonly known as lumpsuckers or lumpfish, in the order Scorpaeniformes. They are found in the cold waters of the Arctic Ocean, Arctic, Atlantic Ocean, North Atlantic, and Pacific Ocean, North Paci ...
s are found in temperate northern waters. They live on the seafloor, and are temporary residents of rocky pools in late winter and early spring when they spawn. The body of the lumpsucker is scaleless and covered with small lumps. They have a large sucking disc on their underside which they use to cling to surfaces. They are normally a blue to slate-grey colour, and are effectively camouflaged to look like stones. They are portly, nearly spherical, poor swimmers, reaching lengths up to . After the female lumpsucker lays eggs, the male takes over, clamping itself to a rock where it guards the eggs. When they hatch, lumpsuckers look like tiny tadpoles. They remain in shallow water and rock pools, hiding amongst seaweed and rocks, until they grow up.


Estuarine fish

Estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
are partly enclosed coastal bodies of water with one or more
river A river is a natural stream of fresh water that flows on land or inside Subterranean river, caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of ...
s or
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a strea ...
s flowing into them, and with a free connection to the open sea.Pritchard, D. W. (1967) ''What is an estuary: physical viewpoint''. p. 3–5 ''in:'' G. H. Lauf (ed.) ''Estuaries'', A.A.A.S. Publ. No. 83, Washington, D.C. These
brackish water Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuary ...
habitats form a transition zone between river environments and ocean environments, and
ecological succession Ecological succession is the process of how species compositions change in an Community (ecology), ecological community over time. The two main categories of ecological succession are primary succession and secondary succession. Primary successi ...
s can form along the way. Estuaries are subject to both marine influences, such as tides, waves, and the influx of saline water; and riverine influences, such as flows of fresh water and sediment. The inflow of both seawater and freshwater provide high levels of nutrients in both the water column and sediment, making estuaries productive natural habitats.McLusky, D.S. and Elliott, M. (2004) "The Estuarine Ecosystem: ecology, threats and management." New York: Oxford University Press Inc. Fishes that spend time in
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
(or
river mouth A river mouth is where a river flows into a larger body of water, such as another river, a lake/reservoir, a bay/gulf, a sea, or an ocean. At the river mouth, sediments are often deposited due to the slowing of the current, reducing the carryin ...
s) need to be
euryhaline Euryhaline organisms are able to adapt to a wide range of salinities. An example of a euryhaline fish is the short-finned molly, '' Poecilia sphenops'', which can live in fresh water, brackish water, or salt water. The green crab ('' Carcinus m ...
(tolerant to a range of salinities). Estuaries provide an unstable environment for fish, where the salinity changes and the waters are often muddy and turbulent. In warmer climates, estuaries have
mangrove A mangrove is a shrub or tree that grows mainly in coastal saline water, saline or brackish water. Mangroves grow in an equatorial climate, typically along coastlines and tidal rivers. They have particular adaptations to take in extra oxygen a ...
s around their edges. At times there may be only a few different fish species present in an estuary, but seasonal migrants, including
eel Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 20 families, 164 genera, and about 1000 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...
s,
salmonid Salmonidae (, ) is a family of ray-finned fish, the only extant member of the suborder Salmonoidei, consisting of 11 extant genera and over 200 species collectively known as "salmonids" or "salmonoids". The family includes salmon (both Atlantic a ...
s, and some
forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish that feed on planktons (i.e. planktivores) and other small aquatic organisms (e.g. krill). They are in turn preyed upon by various predators including larger fish, seabirds ...
such as
herring Herring are various species of forage fish, belonging to the Order (biology), order Clupeiformes. Herring often move in large Shoaling and schooling, schools around fishing banks and near the coast, found particularly in shallow, temperate wate ...
s and
sprat Sprat is the common name applied to a group of forage fish belonging to the genus ''Sprattus'' in the Family (biology), family Clupeidae. The term also is applied to a number of other small sprat-like forage fish (''Clupeoides'', ''Clupeonella ...
s increase the
diversity Diversity, diversify, or diverse may refer to: Business *Diversity (business), the inclusion of people of different identities (ethnicity, gender, age) in the workforce *Diversity marketing, marketing communication targeting diverse customers * ...
in the estuary. River estuaries form important staging points during the migration of anadromous and catadromus fish species, such as
salmon Salmon (; : salmon) are any of several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the genera ''Salmo'' and ''Oncorhynchus'' of the family (biology), family Salmonidae, native ...
and
eel Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 20 families, 164 genera, and about 1000 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...
s, giving them time to form social groups and to adjust to the changes in salinity. Salmon are anadromous, meaning they live in the sea but ascend rivers to spawn; eels are catadromous, living in rivers and streams, but returning to the sea to breed. Besides the species that migrate through estuaries, there are many other fish that use them as "nursery grounds" for spawning or as places young fish can feed and grow before moving elsewhere. For example,
herring Herring are various species of forage fish, belonging to the Order (biology), order Clupeiformes. Herring often move in large Shoaling and schooling, schools around fishing banks and near the coast, found particularly in shallow, temperate wate ...
and
plaice Plaice is a common name for a group of flatfish that comprises four species: the European, American, Alaskan and scale-eye plaice. Commercially, the most important plaice is the European. The principal commercial flatfish in Europe, it is ...
are two commercially important species that use the Thames Estuary for this purpose.
Mangrove swamp Mangrove forests, also called mangrove swamps, mangrove thickets or mangals, are productive wetlands that occur in coastal intertidal zones. Mangrove forests grow mainly at tropical and subtropical latitudes because mangrove trees cannot withs ...
s are associated brackish water habitats. Many, though not all, mangrove swamps fringe estuaries and lagoons where the salinity changes with each tide. Among the most specialised residents of mangrove forests are
mudskippers Mudskippers are any of the 23 extant species of amphibious fish from the subfamily Oxudercinae of the goby family Oxudercidae. They are known for their unusual body shapes, preferences for semiaquatic habitats, limited terrestrial locomotion and j ...
, fish that forage for food on land, and archer fish, perch-like fish that "spit" at insects and other small animals living in the trees, knocking them into the water where they can be eaten. Like estuaries, mangrove swamps are important breeding grounds for many fish, with species such as snappers,
halfbeak Hemiramphidae is a family (biology), family of fishes that are commonly called halfbeaks, spipe fish or spipefish. They are a geographically widespread and numerically abundant family of epipelagic fish inhabiting warm waters around the world. ...
s, and
tarpon Tarpon are fish of the genus ''Megalops''. They are the only members of the family Megalopidae. Of the two species, one (''M. atlanticus'') is native to the Atlantic, and the other (''M. cyprinoides'') to the Indo-Pacific Oceans. Species and ...
spawning or maturing among them.


Coral reef fish

In tropical waters, coral reef fish live amongst or in close relation to
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in group ...
s. Coral reefs form complex
ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...
s with tremendous
biodiversity Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...
. Coral reef fish can be particularly colourful and interesting to watch. Hundreds of species can exist in a small area of a healthy reef, many of them hidden or well camouflaged. Reef fish have developed many ingenious specialisations adapted to survival on the reefs. Coral reefs occupy less than one per cent of the surface area of the world oceans, yet they provide a home for 25 per cent of all marine fish species. Coral reefs often depend on other habitats in the surrounding area for the supply of nutrients, such as
seagrass meadow A seagrass meadow or seagrass bed is an underwater ecosystem formed by seagrasses. Seagrasses are marine (saltwater) plants found in shallow coastal waters and in the brackish waters of estuaries. Seagrasses are flowering plants with stems and ...
s and
mangrove forest Mangrove forests, also called mangrove swamps, mangrove thickets or mangals, are productive wetlands that occur in coastal intertidal zones. Mangrove forests grow mainly at tropical and subtropical latitudes because mangrove trees cannot withsta ...
s. Seagrass and mangroves supply dead plants and animals which are rich in nitrogen and also serve to feed fish and animals from the reef by supplying wood and vegetation. Reefs in turn protect mangroves and seagrass from waves and produce
sediment Sediment is a solid material that is transported to a new location where it is deposited. It occurs naturally and, through the processes of weathering and erosion, is broken down and subsequently sediment transport, transported by the action of ...
for the mangroves and seagrass to root in. Anthias are members of the family
Serranidae Serranidae is a large family (biology), family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, ...
and make up the subfamily Anthiinae. They are widespread in tropical waters. They have been called the "quintessential
reef fish Coral reef fish are fish which live amongst or in close relation to coral reefs. Coral reefs form complex ecosystems with tremendous biodiversity. Among the myriad inhabitants, the fish stand out as colourful and interesting to watch. Hundreds ...
", and make up a sizeable portion of the colourful
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
es seen swarming in
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in group ...
photography. Anthias are mostly small, peaceful, beautiful and popular as
ornamental fish Lists of aquarium life include lists of fish, amphibians, invertebrates and plants in freshwater, brackish and marine aquariums. In fishkeeping, suitable species of aquarium fish, plants and other organisms vary with the size, water chemistry and ...
. They are mainly
zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...
feeders. Anthias shoal and school in large numbers, operating more intimate "harems" within the schools. These harems contain a dominant and colourful male, between 2 and 12 females — who operate a hierarchy among themselves — and one or two "subdominant" males, often less brightly coloured and non-territorial. Within the swarm of females, territorial males perform acrobatic U-swim displays and vigorously defend an area of the reef and its associated harem. Anthias are
protogynous Sequential hermaphroditism (called dichogamy in botany) is one of the two types of hermaphroditism, the other type being simultaneous hermaphroditism. It occurs when the organism's sex changes at some point in its life. A sequential hermaphrodit ...
hermaphrodite A hermaphrodite () is a sexually reproducing organism that produces both male and female gametes. Animal species in which individuals are either male or female are gonochoric, which is the opposite of hermaphroditic. The individuals of many ...
s. All anthias are born female; if a dominant male perishes, the largest female of the group will often change into a male to take its place. This may lead to squabbling between the next largest male and the transforming female, whose hormones are now surging with testosterone. This can turn quite vicious in the limited confines of captivity.
Butterflyfish The butterflyfish are a group of conspicuous tropical ocean, marine fish of the family Chaetodontidae; the bannerfish and coralfish are also included in this group. The approximately 129 species in 12 genera are found mostly on the reefs of the ...
are group of about 120 species belonging to the family Chaetodontidaeof Perchiformes. They include bannerfish and coralfish. They are widespread on coral reefs. Butterflyfish are mostly between in length. The largest species, the
lined butterflyfish The lined butterflyfish (''Chaetodon lineolatus'') is a species of marine ray-finned fish. a butterflyfish belonging to the family Chaetodontidae, one of the largest species in the genus ''Chaetodon''. It has a wide range from the Red Sea to Sou ...
and
saddle butterflyfish The saddle butterflyfish (''Chaetodon ephippium'') is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is found in the Indian and Pacific Oceans from Sri Lanka and the Cocos-Keeling Islands to th ...
, grow to . Many species are brightly coloured and strikingly patterned, though other species are dull in colour. Many have eyespots on their flanks and dark bands across their eyes, not unlike the patterns seen on
butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
wings. Their deep, laterally narrow bodies are easily noticed through the profusion of reef life. The conspicuous colouration of butterflyfish may be intended for interspecies communication. Butterflyfish have uninterrupted
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
s with tail fins that may be rounded or truncated, but are never forked. Generally diurnal and frequenting waters of less than (though some species descend to ), butterflyfish stick to particular home ranges. The
coral Corals are colonial marine invertebrates within the subphylum Anthozoa of the phylum Cnidaria. They typically form compact Colony (biology), colonies of many identical individual polyp (zoology), polyps. Coral species include the important Coral ...
livores are especially territorial, forming mated pairs and staking claim to a specific coral head. Contrastingly, the
zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...
feeders form large conspecific groups. By night butterflyfish hide in reef crevices and exhibit markedly different colouration. Their colouration also makes butterflyfish popular
aquarium An aquarium (: aquariums or aquaria) is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. fishkeeping, Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aquati ...
fish. However, most species feed on
coral Corals are colonial marine invertebrates within the subphylum Anthozoa of the phylum Cnidaria. They typically form compact Colony (biology), colonies of many identical individual polyp (zoology), polyps. Coral species include the important Coral ...
polyps and
sea anemone Sea anemones ( ) are a group of predation, predatory marine invertebrates constituting the order (biology), order Actiniaria. Because of their colourful appearance, they are named after the ''Anemone'', a terrestrial flowering plant. Sea anemone ...
s, which can result in problems for the hobby aquarists.
Clownfish Clownfishes or anemonefishes (genus ''Amphiprion'') are saltwater fishes found in the warm and tropical waters of the Indo-Pacific. They inhabit mainly coral reefs and have a distinctive colouration typically consisting of white vertical bars on ...
, anemonefish and
damselfish Damselfish are those fish within the subfamilies Abudefdufinae, Chrominae, Lepidozyginae, Pomacentrinae, and Stegastinae within the family Pomacentridae. Most species within this group are relatively small, although the four largest speci ...
are among about 360 species classified in the family
Pomacentridae Pomacentridae is a family of ray-finned fish, comprising the damselfishes and clownfishes. This family were formerly placed in the order Perciformes or as indeterminate percomorphs, but are now considered basal blenniiforms. They are primaril ...
. Most Pomacentrids are associated with coral reefs in the Indo-West Pacific, with a few species occurring in temperate waters. Some species are native to freshwater or brackish
estuarine An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
environments. Most live in shallow water, from , although some species are found below . Most species are specialists, living in specific parts of the reef, such as sandy lagoons, steep reef slopes, or areas exposed to strong wave action. In general, the coral is used as shelter, and many species can only survive in its presence. The bottom-dwelling species are territorial, occupying and defending a portion of the reef, often centred around an area of shelter. By keeping away other species of fish, some pomacentrids encourage the growth of thick mats of algae within their territories, leading to the common name farmerfish. Different species display a wide range of colours, although some are relatively drab. Pomacentrids are omnivorous or herbivorous, feeding off algae,
plankton Plankton are the diverse collection of organisms that drift in Hydrosphere, water (or atmosphere, air) but are unable to actively propel themselves against ocean current, currents (or wind). The individual organisms constituting plankton are ca ...
, and small bottom-dwelling
crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
s. A small number eat coral.
Goatfish The goatfishes are ray-finned fish of the family Mullidae, the only family in the suborder Mulloidei of the order Syngnathiformes. The family is also sometimes referred to as the red mullets, which also refers more narrowly to the genus '' Mul ...
es are a family Mullidae of about 55 species of perciform fishes, associated worldwide with tropical
reef A reef is a ridge or shoal of rock, coral, or similar relatively stable material lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic component, abiotic (non-living) processes such as deposition (geol ...
s. They are typically about 20 cm long, though the dash-and-dot goatfish, grows to 55 cm. Goatfish are tireless
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". ...
feeders, possess a pair of long
chemosensory A chemoreceptor, also known as chemosensor, is a specialized sensory receptor which transduces a chemical substance (endogenous or induced) to generate a biological signal. This signal may be in the form of an action potential, if the chemorecept ...
barbels In fish anatomy and turtle anatomy, a barbel is a slender, whisker like sensory organ near the mouth (sometimes called whiskers or tendrils). Fish that have barbels include the catfish, the carp, the goatfish, the hagfish, the sturgeon, the z ...
("whiskers") protruding from their chins resembling a goat's beard. They use these to rifle through the sediments in search of a meal. Like goats, they seek anything edible;
worm Worms are many different distantly related bilateria, bilateral animals that typically have a long cylindrical tube-like body, no limb (anatomy), limbs, and usually no eyes. Worms vary in size from microscopic to over in length for marine ...
s,
crustacea Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
ns,
mollusc Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
s and other small
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s are staples. Many species of goatfish are conspicuously coloured and have the ability to change their colouration depending on their current activity. By day, many form large inactive (non-feeding) schools: these aggregates may contain both conspecifics and heterospecifics. For example, the yellowfin goatfish school with blue-striped snappers. When they do that, the yellowfins changes its colouration to match that of the snapper. By night the schools disperse and individual goatfish head their separate ways to loot the sands. The diurnal goldsaddle goatfish changes from a lemon-yellow to a pale cream when feeding. Other nocturnal feeders will shadow the active goatfish, waiting patiently for overlooked morsels. Goatfish stay within the shallows, going no deeper than about 110 metres. Most species do not tolerate
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
water, so they do not enter
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
or the mouths of
river A river is a natural stream of fresh water that flows on land or inside Subterranean river, caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of ...
s.


Other nearshore fish

Other nearshore or shallow water fish live near the shore in depths of less than 10 metres. They occupy the areas over sandy or rocky bottoms, and can be associated with
seagrass meadow A seagrass meadow or seagrass bed is an underwater ecosystem formed by seagrasses. Seagrasses are marine (saltwater) plants found in shallow coastal waters and in the brackish waters of estuaries. Seagrasses are flowering plants with stems and ...
s and
kelp forest Kelp forests are underwater areas with a high density of kelp, which covers a large part of the world's coastlines. Smaller areas of anchored kelp are called kelp beds. They are recognized as one of the most productive and dynamic ecosystems on E ...
s. They can be divided into
demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...
and
pelagic fish Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that live on or near the bottom, and reef fish that are associated with coral reefs. ...
. Demersal fish live on or near the sea floor, while pelagic fish live in the
water column The (oceanic) water column is a concept used in oceanography to describe the physical (temperature, salinity, light penetration) and chemical ( pH, dissolved oxygen, nutrient salts) characteristics of seawater at different depths for a defined ...
away the sea floor. Examples of such shallow water demersal fish, found in both tropical and temperate waters around the world, are seahorses, triplefins, wrasse and flounder. As demersal fish, all these fish spend most of their time on or near the sea floor. *
Flatfish A flatfish is a member of the Ray-finned fish, ray-finned demersal fish Order (biology), suborder Pleuronectoidei, also called the Heterosomata. In many species, both eyes lie on one side of the head, one or the other migrating through or around ...
are superbly adapted
groundfish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...
, found on muddy and sandy sea floors. In many species both eyes lie on one side of the head, one or the other migrating through and around the head during development. Some species face their "left" side upward, some face their "right" side upward, and others face either side upward. Some flatfish can camouflage themselves on the ocean floor. *
Wrasse The wrasses are a family, Labridae, of marine ray-finned fish, many of which are brightly colored. The family is large and diverse, with over 600 species in 81 genera, which are divided into nine subgroups or tribes. They are typically small, ...
are a large family of mainly small fish, usually less than long. Most wrasse are loners that prefer habitats such as coral reefs and rocky shores. They live close to the substrate, eating small invertebrates and almost anything else that lurks on the bottom. Many are brightly coloured. They have thick lips and use their sharp teeth to pick small creatures off the rocks. Many smaller wrasses follow the feeding trails of larger fish, picking up invertebrates disturbed by their passing. * Triplefins are a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of fish. They are usually found around
coral reefs A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. ...
and rocks, usually in shallow, clear sunlit waters such as lagoons and seaward reefs. Triplefins have three
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
s (hence the name). They are small fish, usually less than six cm long. Brightly coloured, often for reasons of camouflage, they are nervous and retreat to rock crevices at any perceived threat. *
Seahorse A seahorse (also written ''sea-horse'' and ''sea horse'') is any of 46 species of small marine Osteichthyes, bony fish in the genus ''Hippocampus''. The genus name comes from the Ancient Greek (), itself from () meaning "horse" and () meanin ...
s are a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of fish. They prefer sheltered harbours, estuaries and other shallow coastal waters, where they hunt tiny crustaceans. They bob around in sheltered areas such as
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in group ...
s, mangrove stands and
seagrass meadow A seagrass meadow or seagrass bed is an underwater ecosystem formed by seagrasses. Seagrasses are marine (saltwater) plants found in shallow coastal waters and in the brackish waters of estuaries. Seagrasses are flowering plants with stems and ...
s and
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
. They are camouflaged with murky patterns that blend into kelp and sea grass backgrounds. During social moments or in unusual surroundings, seahorses can turn on bright colours. File:Hippocampus.jpg, Seahorse File:Pseudopleuronectes americanus.jpg, Flatfish File:Cleaning station.JPG, Cleaner wrasses working on a dragon wrasse File:Common triplefin.png, Common triplefin Examples of shallow water pelagic fish, found in both tropical and temperate waters around the world, are grey mullet, sprats and garfish. As pelagic fish, all these fish spend most of their time living in the
water column The (oceanic) water column is a concept used in oceanography to describe the physical (temperature, salinity, light penetration) and chemical ( pH, dissolved oxygen, nutrient salts) characteristics of seawater at different depths for a defined ...
away the sea floor. * The grey mullet are medium size fish, typically about long. They are often caught with
seine net Seine fishing (or seine-haul fishing; ) is a method of fishing that employs a surrounding net, called a seine, that hangs vertically in the water with its bottom edge held down by weights and its top edge buoyed by floats. Seine nets can be de ...
s. * The
garfish The garfish (''Belone belone''), also known as the garpike, needlefish or sea needle, is a pelagic, oceanodromous needlefish found in brackish and marine waters of the Atlantic Ocean and the Mediterranean, Caribbean, Black, and Baltic Seas. De ...
is a long, slender fish, looking like a spear, which feeds on seagrass fragments, shrimps and crab larvae. In turn it is preyed on by larger fish and, since it is often near the surface,
cormorant Phalacrocoracidae is a family of approximately 40 species of aquatic birds commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the International Ornithologists' Union (IOU) ado ...
s and
gannet Gannets are seabirds comprising the genus ''Morus'' in the family Sulidae, closely related to boobies. They are known as 'solan' or 'solan goose' in Scotland. A common misconception is that the Scottish name is 'guga' but this is the Gaelic n ...
s. File:Mucep u0.gif, Grey mullet File:Belone belone1.jpg,
Garfish The garfish (''Belone belone''), also known as the garpike, needlefish or sea needle, is a pelagic, oceanodromous needlefish found in brackish and marine waters of the Atlantic Ocean and the Mediterranean, Caribbean, Black, and Baltic Seas. De ...


Coastal pelagic fish


Plankton feeding

At the base of
food chain A food chain is a linear network of links in a food web, often starting with an autotroph (such as grass or algae), also called a producer, and typically ending at an apex predator (such as grizzly bears or killer whales), detritivore (such as ...
s are the
primary producer Primary or primaries may refer to: Arts, entertainment, and media Music Groups and labels * Primary (band), from Australia * Primary (musician), hip hop musician and record producer from South Korea * Primary Music, Israeli record label Work ...
s. In the ocean these primary producers are mainly a type of
plankton Plankton are the diverse collection of organisms that drift in Hydrosphere, water (or atmosphere, air) but are unable to actively propel themselves against ocean current, currents (or wind). The individual organisms constituting plankton are ca ...
, microscopic
phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater Aquatic ecosystem, ecosystems. The name comes from the Greek language, Greek words (), meaning 'plant', and (), mea ...
which drift in the
water column The (oceanic) water column is a concept used in oceanography to describe the physical (temperature, salinity, light penetration) and chemical ( pH, dissolved oxygen, nutrient salts) characteristics of seawater at different depths for a defined ...
. Phytoplankton need sunlight for
photosynthesis Photosynthesis ( ) is a system of biological processes by which photosynthetic organisms, such as most plants, algae, and cyanobacteria, convert light energy, typically from sunlight, into the chemical energy necessary to fuel their metabo ...
to power
carbon fixation Biological carbon fixation, or сarbon assimilation, is the Biological process, process by which living organisms convert Total inorganic carbon, inorganic carbon (particularly carbon dioxide, ) to Organic compound, organic compounds. These o ...
, so they are mainly located in sunlit surface waters. Phytoplankton also need and rapidly use nutrients in the water column.Anderson, G (2003
Coral Reef Formation
''Marine Science''.
The phytoplankton are eaten by
zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...
, which in turn are eaten by predatory zooplankton.
Filter feeder Filter feeders are aquatic animals that acquire nutrients by feeding on organic matters, food particles or smaller organisms (bacteria, microalgae and zooplanktons) suspended in water, typically by having the water pass over or through a s ...
s then eat the plankton and larger
predatory fish Predatory fish are hypercarnivorous fish that actively prey upon other fish or aquatic animals, with examples including shark, billfish, barracuda, alligator gar, tuna, dolphinfish, walleye, perch and salmon. Some omnivorous fish, such as t ...
eat the filter feeders (see diagram at the right).Mann, K.H., Lazier, J.R.N. (2006) ''Dynamics of Marine Ecosystems: Biological-Physical Interactions in the Oceans''. Oxford: Blackwell Publishing Ltd. Most filter-feeding pelagic fish found in coastal waters are small, silvery
forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish that feed on planktons (i.e. planktivores) and other small aquatic organisms (e.g. krill). They are in turn preyed upon by various predators including larger fish, seabirds ...
. Forage fish include fishes of the family
Clupeidae Clupeidae is a family of clupeiform ray-finned fishes, comprising, for instance, the herrings and sprats. Many members of the family have a body protected with shiny cycloid (very smooth and uniform) scales, a single dorsal fin, and a fusiform ...
(
herring Herring are various species of forage fish, belonging to the Order (biology), order Clupeiformes. Herring often move in large Shoaling and schooling, schools around fishing banks and near the coast, found particularly in shallow, temperate wate ...
s,
shad The Alosidae, or the shads, are a family (biology), family of clupeiform fishes. The family currently comprises four genera worldwide, and about 32 species. The shads are Pelagic fish, pelagic (open water) schooling fish, of which many are anadr ...
,
sardine Sardine and pilchard are common names for various species of small, oily forage fish in the herring suborder Clupeoidei. The term "sardine" was first used in English during the early 15th century; a somewhat dubious etymology says it com ...
s and
pilchard Sardine and pilchard are common names for various species of small, oily forage fish in the herring suborder Clupeoidei. The term "sardine" was first used in English during the early 15th century; a somewhat dubious etymology says it comes ...
s,
hilsa The ilish (''Tenualosa ilisha'') (), also known as the ilishi, hilsa, hilsa herring or hilsa shad, is a species of fish related to the herring, in the family Clupeidae. It is a very popular and sought-after food in the Bengal region, and is ...
,
menhaden Menhaden, also known as mossbunker, bunker, and "the most important fish in the sea", are forage fish of the genera ''Brevoortia'' and ''Ethmidium'', two genera of marine fish in the order Clupeiformes. ''Menhaden'' is a blend of ''poghaden'' ...
and
sprat Sprat is the common name applied to a group of forage fish belonging to the genus ''Sprattus'' in the Family (biology), family Clupeidae. The term also is applied to a number of other small sprat-like forage fish (''Clupeoides'', ''Clupeonella ...
s), as well as
anchovies An anchovy is a small, common forage fish of the family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 species are placed in 1 ...
,
capelin The capelin or caplin (''Mallotus villosus'') is a small forage fish of the smelt family found in the North Atlantic, North Pacific and Arctic oceans. In summer, it grazes on dense swarms of plankton at the edge of the ice shelf. Larger capel ...
and
halfbeak Hemiramphidae is a family (biology), family of fishes that are commonly called halfbeaks, spipe fish or spipefish. They are a geographically widespread and numerically abundant family of epipelagic fish inhabiting warm waters around the world. ...
s. They use
schooling A school is the educational institution (and, in the case of in-person learning, the building) designed to provide learning environments for the teaching of students, usually under the direction of teachers. Most countries have systems of fo ...
strategies to avoid predators, and different schools of forage fish often associate with each other in open coastal waters. Forage fish feed near the base of the food chain on
plankton Plankton are the diverse collection of organisms that drift in Hydrosphere, water (or atmosphere, air) but are unable to actively propel themselves against ocean current, currents (or wind). The individual organisms constituting plankton are ca ...
and fry (recently hatched fish), often by
filter feeding Filter feeders are aquatic animals that acquire nutrients by feeding on organic matters, food particles or smaller organisms (bacteria, microalgae and zooplanktons) suspended in water, typically by having the water pass over or through a spe ...
. In turn, they are preyed on by larger predatory fish,
seabird Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine ecosystem, marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ...
s and
marine mammal Marine mammals are mammals that rely on marine ecosystems for their existence. They include animals such as cetaceans, pinnipeds, sirenians, sea otters and polar bears. They are an informal group, unified only by their reliance on marine enviro ...
s. Worldwide, there are five major coastal currents associated with upwelling areas: the
Canary Current The Canary Current is a wind-driven surface current that is part of the North Atlantic Gyre. This eastern boundary current branches south from the North Atlantic Current and flows southwest about as far as Senegal where it turns west and later jo ...
(off
Northwest Africa The Maghreb (; ), also known as the Arab Maghreb () and Northwest Africa, is the western part of the Arab world. The region comprises western and central North Africa, including Algeria, Libya, Mauritania, Morocco, and Tunisia. The Maghreb al ...
), the
Benguela Current The Benguela Current is the broad, northward flowing ocean current that forms the eastern portion of the South Atlantic Ocean gyre. The current extends from roughly Cape Point in the south, to the position of the Angola-Benguela Front in the no ...
(off
southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
), the
California Current The California Current () is a cold water Pacific Ocean ocean current, current that moves southward along the western coast of North America, beginning off southern British Columbia and ending off southern Baja California Sur. It is considered an ...
(off
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
and
Oregon Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while t ...
), the
Humboldt Current The Humboldt Current, also called the Peru Current, is a cold, low-salinity ocean current that flows north along the western coast of South America.Montecino, Vivian, and Carina B. Lange. "The Humboldt Current System: Ecosystem components and pro ...
(off
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
and
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
), and the Somali Current (off
Western India Western India is a loosely defined region of India consisting of western states of India, Republic of India. The Ministry of Home Affairs (India), Ministry of Home Affairs in its Western Zonal Council Administrative divisions of India, Adminis ...
). All of these currents support major fisheries. Many forage fish are important commercial species, and the schools can be targeted by spotter planes. The fish are caught by
purse seiner A fishing vessel is a boat or ship used to fishing, catch fish and other valuable nektonic aquatic animals (e.g. shrimps/prawns, krills, coleoids, etc.) in the sea, lake or river. Humans have used different kinds of surface vessels in commercial ...
s—fishing boats that use nets to enclose the fish—and can be
overfished Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in the ...
. File:Anchovy closeup.jpg,
Schooling A school is the educational institution (and, in the case of in-person learning, the building) designed to provide learning environments for the teaching of students, usually under the direction of teachers. Most countries have systems of fo ...
anchovies An anchovy is a small, common forage fish of the family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 species are placed in 1 ...
File:Clupea sprattus.jpg, European sprat File:Herring2.jpg,
Atlantic herring Atlantic herring (''Clupea harengus'') is a herring in the family Clupeidae. It is one of the most abundant fish species in the world. Atlantic herrings can be found on both sides of the northern Atlantic Ocean, congregating in large schools. ...
File:Heringsschwarm.gif, Underwater video loop of a school of herrings migrating to their spawning grounds in the
Baltic Sea The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by the countries of Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden, and the North European Plain, North and Central European Plain regions. It is the ...


Predatory

Predatory pelagic fishes found on continental shelves worldwide in both tropical and temperate waters include porgies,
barracuda A barracuda is a large, predatory, ray-finned, saltwater fish of the genus ''Sphyraena'', the only genus in the family Sphyraenidae, which was named by Constantine Samuel Rafinesque in 1815. It is found in tropical and subtropical oceans worldw ...
,
amberjack Amberjacks are Atlantic and Pacific Ocean, Pacific fish in the genus ''Seriola'' of the family Carangidae. They are widely consumed across the world in various cultures, most notably for Pacific amberjacks in Japanese cuisine; they are most oft ...
s and
cutlassfish The cutlassfishes are about 45 species of predatory ray-finned fish in the family Trichiuridae of the order Scombriformes found in seas throughout the world. Fish of this family are long, slender, and generally steely blue or silver in colour, ...
es. They tend to be larger fish, and are carnivorous, feeding on the smaller, silvery
forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish that feed on planktons (i.e. planktivores) and other small aquatic organisms (e.g. krill). They are in turn preyed upon by various predators including larger fish, seabirds ...
that eat plankton (see section above). Some species also feed on crabs and other invertebrates, foraged from the sea floor. *
Mackerel Mackerel is a common name applied to a number of different species of pelagic fish, mostly from the family Scombridae. They are found in both temperate and tropical seas, mostly living along the coast or offshore in the oceanic environment. ...
scom* * Porgies sometimes called sea breams, are any of about 100 species belonging to the family Sparidae. Porgies usually have high backs and a single dorsal fin, like snapper or grunt fishes (grunts are named for the sound they make grinding their teeth). They are bottom feeding pelagic fishes, with small mouths equipped with strong teeth adapted for handling small fishes and invertebrates with hard shells. Most do not exceed a size of about , but some may grow to four times that length. They often school, and will migrate between reefs. Larger fish enter estuaries and harbours. *
Barracuda A barracuda is a large, predatory, ray-finned, saltwater fish of the genus ''Sphyraena'', the only genus in the family Sphyraenidae, which was named by Constantine Samuel Rafinesque in 1815. It is found in tropical and subtropical oceans worldw ...
have long slender bodies typically about long. They have a wicked set of teeth and are ferocious predators. They feed on
crustaceans Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of Arthropod, arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquat ...
,
cephalopods A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symm ...
and small fish like
anchovy An anchovy is a small, common forage fish of the Family (biology), family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 speci ...
and
pilchard Sardine and pilchard are common names for various species of small, oily forage fish in the herring suborder Clupeoidei. The term "sardine" was first used in English during the early 15th century; a somewhat dubious etymology says it comes ...
. Barracouta often hunt in schools near the bottom or midwater, and sometimes even near the surface at night. File:Calamus bajonado.jpg, The jolthead porgy is a porgi gruntPollnac, R.B. 1998 Aspects of the human ecology of the coral reefs of Discovery Bay. p. 141-199. In R.B. Pollnac (ed.) Rapid assessment of management parameters for coral reefs. Coastal Resources Center, University of Rhode Island. CMR 2205, ICLARM 1445. File:Great Barracuda off the Netherland Antilles.jpg, Great barracuda and jacks File:Seriolalalandii.jpg, The yellowtail amberjack, pound for pound, is one of the hardest fighting fish in the ocean. File:Atlantic_cutlassfish.jpg, The
largehead hairtail The largehead hairtail (''Trichiurus lepturus'') or beltfish is a member of the cutlassfish family, Trichiuridae. This common to abundant species is found in tropical and temperate oceans throughout the world. The taxonomy is not fully resolved, ...
is a
cutlassfish The cutlassfishes are about 45 species of predatory ray-finned fish in the family Trichiuridae of the order Scombriformes found in seas throughout the world. Fish of this family are long, slender, and generally steely blue or silver in colour, ...
*
Cutlassfish The cutlassfishes are about 45 species of predatory ray-finned fish in the family Trichiuridae of the order Scombriformes found in seas throughout the world. Fish of this family are long, slender, and generally steely blue or silver in colour, ...
es are a group of about 40 species belonging to the family Trichiuridae. They are ocean fish which regularly stray into coastal waters around the world. Fish of this family are long, slender, and generally steely blue or silver in colour, giving rise to their name. They have reduced or absent
pelvic The pelvis (: pelves or pelvises) is the lower part of an anatomical trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis or pelvic skeleton). ...
and
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
s, giving them an
eel Eels are ray-finned fish belonging to the order Anguilliformes (), which consists of eight suborders, 20 families, 164 genera, and about 1000 species. Eels undergo considerable development from the early larval stage to the eventual adult stage ...
-like appearance, and large fang-like teeth. * Jacks,
amberjack Amberjacks are Atlantic and Pacific Ocean, Pacific fish in the genus ''Seriola'' of the family Carangidae. They are widely consumed across the world in various cultures, most notably for Pacific amberjacks in Japanese cuisine; they are most oft ...
s,
pompano Pompanos ( ) are saltwater fish, marine fish in the genus ''Trachinotus'' in the family (biology), family Carangidae (better known as "slabs"). Pompano may also refer to various other, similarly shaped members of the Carangidae, or the Order (bi ...
s, horse mackerel, scads, leatherjackets and
trevally The Carangidae are a family of ray-finned fish that includes the jacks, pompanos, jack mackerels, runners, trevallies, and scads. It is the largest of the six families included within the order Carangiformes. Some authorities classify it as the ...
are fish of the family ''
Carangidae The Carangidae are a family of ray-finned fish that includes the jacks, pompanos, jack mackerels, runners, trevallies, and scads. It is the largest of the six families included within the order Carangiformes. Some authorities classify it as the ...
''. Found in most coastal waters, they are fast predatory fishes that hunt in the waters above
reef A reef is a ridge or shoal of rock, coral, or similar relatively stable material lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic component, abiotic (non-living) processes such as deposition (geol ...
s and in the open sea; some dig in the sea floor for invertebrates (some can also filter feed, such as the white trevally). The largest fish in the family, the
giant trevally The giant trevally (''Caranx ignobilis''), also known as the lowly trevally, barrier trevally, ronin jack, giant kingfish, or ''ulua'', is a species of large ocean, marine fish classified in the jack Family (biology), family, Carangidae. The gian ...
, grows up to 1.7 m in length; most fish in the family reach a maximum length of 25–100 cm. The family contains many important commercial and game fish, notably the Pacific jack mackerel and the other jack mackerels in the genus '' Trachurus''. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
of this genus is the
Atlantic horse mackerel The Atlantic horse mackerel (''Trachurus trachurus''), also known as the European horse mackerel or common scad, is a species of jack mackerel in the family Carangidae, which includes the jacks, pompanos and trevallies. It is found in the easte ...
. Jack mackerels are an important inshore commercial species. **
Amberjack Amberjacks are Atlantic and Pacific Ocean, Pacific fish in the genus ''Seriola'' of the family Carangidae. They are widely consumed across the world in various cultures, most notably for Pacific amberjacks in Japanese cuisine; they are most oft ...
s are a group of nine species belonging to the genus ''Seriola'' within the family Carangidae. Mainly open water fish, they can follow small
forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish that feed on planktons (i.e. planktivores) and other small aquatic organisms (e.g. krill). They are in turn preyed upon by various predators including larger fish, seabirds ...
into estuaries and enclosed waters, where they will also hunt for crustaceans. Amberjacks are fast swimming and aggressive predators that often hunt in schools around offshore reefs. The yellowtail amberjack can reach 1.8 m in length and weigh 60 kilograms.


Demersal pelagic fish

Fish that live on or in close association with the sea floor are called
demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...
. This section discusses the coastal demersal fish that live on the continental shelf, but are living further from the coast and in deeper water than the nearshore fish discussed above. Demersal fish are white fish. Unlike
oily fish Oily fish are fish species with fish oil, oil (fats) in soft tissues and in the coelomic cavity around the Gut (zoology), gut. Their fillet (cut), fillets may contain up to 30% oil, although this figure varies both within and between species. ...
, white fish contain
oils An oil is any nonpolar chemical substance that is composed primarily of hydrocarbons and is hydrophobic (does not mix with water) and lipophilic (mixes with other oils). Oils are usually flammable and surface active. Most oils are unsaturat ...
only in their
liver The liver is a major metabolic organ (anatomy), organ exclusively found in vertebrates, which performs many essential biological Function (biology), functions such as detoxification of the organism, and the Protein biosynthesis, synthesis of var ...
, rather than in the gut, and can therefore be gutted as soon as they are caught, on board the ship. White fish has dry and white flesh. They can be divided into
benthopelagic fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...
(mostly "round" fish) which live ''near'' the sea bed, such as
cod Cod (: cod) is the common name for the demersal fish genus ''Gadus'', belonging to the family (biology), family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gad ...
, and
benthic fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...
(
flatfish A flatfish is a member of the Ray-finned fish, ray-finned demersal fish Order (biology), suborder Pleuronectoidei, also called the Heterosomata. In many species, both eyes lie on one side of the head, one or the other migrating through or around ...
) such as
plaice Plaice is a common name for a group of flatfish that comprises four species: the European, American, Alaskan and scale-eye plaice. Commercially, the most important plaice is the European. The principal commercial flatfish in Europe, it is ...
which live ''on'' the sea bed. Benthic fish tend to be "flat", so they can lie on the bottom. *
Cod Cod (: cod) is the common name for the demersal fish genus ''Gadus'', belonging to the family (biology), family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gad ...
-like fishes are a number of round benthopelagic species belonging to the order
Gadiformes Gadiformes , also called the Anacanthini, are an order of ray-finned fish that include the cod, hakes, pollock, haddock, burbot, rocklings and moras, many of which are food fish of major commercial value. They are mostly marine fish found thr ...
, such as
Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...
and
Pacific cod The Pacific cod (''Gadus macrocephalus)'' is a species of ray-finned fish in the family Gadidae. It is a bottom-dwelling fish found in the northern Pacific Ocean, mainly on the continental shelf and upper slopes, to depths of about . It can grow ...
,
morid cod The Moridae are a family of cod-like fishes, known as codlings, hakelings, and moras. Morids are marine fishes found throughout the world, and may be found at depths to , although most prefer shallower waters. In appearance, they greatly resemb ...
,
haddock The haddock (''Melanogrammus aeglefinus'') is a saltwater ray-finned fish from the Family (biology), family Gadidae, the true cods. It is the only species in the Monotypy, monotypic genus ''Melanogrammus''. It is found in the North Atlantic Oce ...
and
pollock Pollock or pollack (pronounced ) is the common name used for either of the two species of North Atlantic ocean, marine fish in the genus ''Pollachius''. ''Pollachius pollachius'' is referred to as "pollock" in North America, Ireland and the Unit ...
, including the highly commercial
Alaska pollock The Alaska pollock or walleye pollock (''Gadus chalcogrammus'') is a marine fish species of the cod genus ''Gadus'' and family Gadidae. It is a semi-pelagic Shoaling and schooling, schooling fish widely distributed in the North Pacific Ocean, No ...
. Cod-like fishes are often found in large schools over sandy or muddy bottoms. They have a barbel (fleshy filament) on their lower jaw which they use to detect prey buried in the sand or mud. Some migrate to warm water in winter to spawn. * John Dory are fishes of the genus ''Zeus''. They have a widespread distribution and are typically found near the seabed in depths from . The John Dory grows to a maximum length of . Although it is a benthopelagic fish, its body is flat and it can hardly be seen from the front because it is so thin. It is a poor swimmer with long spines on the dorsal fin. It has a large dark eyespot on the side of its body which is used to confuse prey, which are scooped up in its big mouth. Large eyes at the front of the head provide it with bifocal vision and depth perception, which are important for predators. The John Dory usually gets its food by stalking it then shooting out a tube in its mouth to capture its prey. It eats forage fish, and occasionally
squid A squid (: squid) is a mollusc with an elongated soft body, large eyes, eight cephalopod limb, arms, and two tentacles in the orders Myopsida, Oegopsida, and Bathyteuthida (though many other molluscs within the broader Neocoleoidea are also ...
and cuttlefish. In turn, they are preyed on by sharks, like the dusky shark, and other large bony fish. They are normally solitary. * Turbot and Brill (fish), brill are benthic flatfish, resembling flounder and sole, but found in deeper offshore waters on the continental shelf. They are brownish-green, with dark blotches on the turbot and mottling on the brill. They are fished by coastal trawlers. File:Boreogadus saida.jpg, The polar cod is found further north than any other fish species. It frequents river mouths and feeds on
plankton Plankton are the diverse collection of organisms that drift in Hydrosphere, water (or atmosphere, air) but are unable to actively propel themselves against ocean current, currents (or wind). The individual organisms constituting plankton are ca ...
and krill. It is preyed on by narwhals, Beluga (whale), belugas, ringed seals and
seabird Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine ecosystem, marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ...
s. File:MacGillivray, William John Dory.jpg, The John Dory is so thin it can hardly be seen from the front. The large eyespot on the side of its body confuses its prey. File:Psetta_maxima_Luc_Viatour.jpg, The turbot is a large left-eyed
flatfish A flatfish is a member of the Ray-finned fish, ray-finned demersal fish Order (biology), suborder Pleuronectoidei, also called the Heterosomata. In many species, both eyes lie on one side of the head, one or the other migrating through or around ...
usually found not too far from shore in sandy shallow waters. It is a prized food fish. File:Helidonichthys_spinosus.jpg, The red gurnard is a Scorpaeniformes, mail-cheeked fish found to depths of around 180 metres. They often grunt when captured as air is expelled from its air bladder.
* Scorpaeniformes, Mail-cheeked fishes belong to a group of about 30 species in the order Scorpaeniformes. Mail-cheeked fishes are named after a plate of bone that runs across each cheek. They are widespread in all the oceans of the world. Mail-cheeked fishes are carnivore, carnivorous, mostly feeding on crustaceans, such as crabs and shrimp, and on smaller fish. Most species live on the sea bottom in relatively shallow waters, although species are known from mid and deep water, from the mid-water, and even from fresh water. They typically have spiny heads, and rounded pectoral fin, pectoral and
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
s. Most species are less than in length, but the full size range of the order varies from the velvetfishes, which can be just long as adults, to the Lingcod, which can reach in length. ** Chelidonichthys kumu, Red gurnard are mail-cheeked fish. They use their large pectoral fins to rest on the bottom and to detect food. Stargazer (fish), Stargazers are about 50 species of fishes, belonging to the family Uranoscopidae, and found worldwide in shallow waters. Stargazers are venom (poison), venomous; they have two large poison spines situated behind the opercle and above the pectoral fins. They can also deliver electric shocks. They are ambush predators with eyes on top of their heads (thus the name). Stargazers also have a large upward-facing mouth in a large head. They bury themselves in sand with only their eyes showing, and leap upwards to ambush fish and invertebrates overhead. Some species have a worm-shaped lure growing out of the floor of the mouth, which they wiggle to attract prey's attention. Lengths range from 18 Centimetre, cm up to 90 cm, for the giant stargazer ''Kathetostoma giganteum''. Stargazers are a delicacy in some cultures. The venom is destroyed when it is cooked, and stargazers are sold in some fish markets with their electric organ removed. They have been called "the meanest things in creation" and the "worst pet on earth".Grady, Denis
Venom Runs Thick in Fish Families, Researchers Learn
''The New York Times'' 22 August 2006.
* Sandperches are a family, Pinguipedidae, containing 63 species of fishes in the order Perciformes. They are
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". ...
carnivores, feeding on small fish and invertebrates. Examples are the redbanded weever, yellow weaver (fish), yellow weaver and blue cod. They are often caught in pots like crayfish. * Medusa fishes are a family Centrolophidae of 31 species of perciform fishes. They are found in temperate and tropical waters throughout the world, usually feeding on fish, crustaceans and small squid near rough sea floors on continental shelf and slope. Examples are Hyperoglyphe, barrelfish, southern driftfish, imperial blackfish, the Japanese butterfish, Japanese and pelagic butterfish, the New Zealand ruffe, New Zealand and Tasmanian ruffe, and the Common warehou, common, Silver warehou, silver and white warehou. The young of some species associate with jellyfish, which provides them with protection from predators and opportunities to scavenge the remains of the jellyfish's meals. The young of other species associate with large masses of floating kelp. File:Parapercis hexophtalma 1.jpg, A speckled sand perch perched on coral sand File:Centrolophus niger.jpg, The rudderfish (''Centrolophus niger'') is a medusa fish File:Epinephelus malabaricus.jpg, Malabar grouper File:Polyprion americanus.2 - Aquarium Finisterrae.JPG, Atlantic wreckfish inhabit caves and wrecks. They are good game fish, reaching a maximum reported length of and weight of . * Grouper are fish belonging to a number of genus, genera in the subfamily Epinephelinae of the family
Serranidae Serranidae is a large family (biology), family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, ...
, in the order Perciformes. Species of grouper include the Black grouper, black, Comet grouper, comet, Gag grouper, gag, Giant grouper, giant, Atlantic goliath grouper, Goliath, Nassau grouper, Nassau, Saddletail grouper, saddletail, Tiger grouper, tiger, Warsaw grouper, Warsaw, White grouper, white and yellowfin grouper. Typical lengths are 80–120 centimetres. They inhabit depths from reefs near the surface down to over 400 metres. They feed on just about any moving animal they encounter. Groper are important inshore commercial fish, usually caught with gill nets (in earlier times longline fishing, longlines were used). * Wreckfish are a family Polyprionidae of perciform fishes, found on the floor of the continental shelf and slope where they inhabit caves and shipwrecks (thus their common name). The Atlantic wreckfish is at depths between . They are largely a solitary fish, though juveniles school below floating objects. Their diet includes large ocean cephalopods, crustaceans, and other bottom-dwelling fishes.


See also

* Coral reef fish * Demersal fish * Forage fish * Marine habitats * Moorish idol * Monkfish * NaGISA * Pelagic fish * Shoaling and schooling * Weever


Notes


References

* Elliott, Michael and Hemingway, Krystal (2002
''Fishes in estuaries''
John Wiley & Sons. . * Humann, Paul and DeLoach, Ned (2008
''Coastal Fish Identification: California to Alaska''
New World Publications Incorporated. . * Moyle, PB and Cech, JJ (2004
''Fishes, An Introduction to Ichthyology''
5th Ed, Benjamin Cummings. * Hinrichsen D (1999
''Coastal Waters of the World: Trends, Threats, and Strategies''
Island Press. * Mann KH (2000
''Ecology of coastal waters: with implications for management''
(Vol 8 of ''Studies in ecology'') Wiley-Blackwell.


External links

* Walrond, Carl
"Coastal fish"
– ''Te Ara: The Encyclopedia of New Zealand'' (updated 23 November 2009)
Marine Biology 2: Water layering and water movements


Retrieved 20 February 2010.
Estuarine Fish Kills in Morth Carolina
Retrieved 20 February 2010.

{{Fishery science topics Marine fish