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''Equisetum hyemale'' (rough horsetail) is an
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of ...
pteridophyte A pteridophyte is a vascular plant (with xylem and phloem) that reproduces by means of spores. Because pteridophytes produce neither flowers nor seeds, they are sometimes referred to as " cryptogams", meaning that their means of reproduction is ...
in the horsetail family
Equisetaceae Equisetaceae, also known as the horsetail family, is a family of ferns and the only surviving family of the order Equisetales, with one surviving genus, ''Equisetum'', comprising about twenty species. Evolution and systematics Equisetaceae is ...
native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...
to
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
and
Greenland Greenland is an autonomous territory in the Danish Realm, Kingdom of Denmark. It is by far the largest geographically of three constituent parts of the kingdom; the other two are metropolitan Denmark and the Faroe Islands. Citizens of Greenlan ...
. It was formerly widely treated in a broader sense including a
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
(subsp. ''affine'') in North America, but this is now treated as a separate species, '' Equisetum praealtum''.


Distribution

''Equisetum hyemale'' is native to central and northern Eurasia, including Iceland, Greenland, Kamchatka and Japan, where it forms
clonal colonies A clonal colony or genet is a group of genetically identical individuals, such as plants, fungi, or bacteria, that have grown in a given location, all originating vegetatively, not sexually, from a single ancestor. In plants, an individual in ...
in mesic (reliably moist) habitats, often in heavy clay or sandy soils in
riparian zone A riparian zone or riparian area is the interface between land and a river or stream. In some regions, the terms riparian woodland, riparian forest, riparian buffer zone, riparian corridor, and riparian strip are used to characterize a ripari ...
s of rivers and streams where it can withstand occasional flooding, but also in lime-rich upland
flushes Flushing is to become markedly red in the face and often other areas of the skin, from various physiological conditions. Flushing is generally distinguished from blushing, since blushing is psychosomatic, milder, generally restricted to the face ...
and
seep A seep or flush is a moist or wet place where water, usually groundwater, reaches the Earth's surface from an underground aquifer. Description Seeps are usually not of sufficient volume to be flowing beyond their immediate above-ground location. ...
s, and can indicate their presence when not flowing.Hyde, H. A., Wade, A. E., & Harrison, S. G. (1978). ''Welsh Ferns''. National Museum of Wales . Other
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s include moist forest and woodland openings, lake and pond shores, ditches,
marsh In ecology, a marsh is a wetland that is dominated by herbaceous plants rather than by woody plants.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p More in genera ...
es and
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s. It grows from between sea level to in elevation. It has been introduced into
Tasmania Tasmania (; palawa kani: ''Lutruwita'') is an island States and territories of Australia, state of Australia. It is located to the south of the Mainland Australia, Australian mainland, and is separated from it by the Bass Strait. The sta ...
.


Description

''Equisetum hyemale'' has vertical jointed reed-like stalks of medium to dark green. The hollow stems are up to in height, and are rarely branched. The stems are 3–6 mm thick with 10–30 conspicuous ridges, which are impregnated with silica and rough to the touch (from which the English name derives). The tiny leaves are joined around the stem, forming a narrow black-green band or sheath at each joint. Like other pteridophytes, the plant reproduces by
spore In biology, a spore is a unit of sexual reproduction, sexual (in fungi) or asexual reproduction that may be adapted for biological dispersal, dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores fo ...
s and does not produce flowers or seeds. The stems remain green during winter in warmer climates, but are generally deciduous in cold climates. It forms dense spreading colonies in full to partial sun.


Taxonomy

Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
was the first to describe rough horsetail with the
binomial Binomial may refer to: In mathematics *Binomial (polynomial), a polynomial with two terms *Binomial coefficient, numbers appearing in the expansions of powers of binomials *Binomial QMF, a perfect-reconstruction orthogonal wavelet decomposition * ...
''Equisetum hyemale'' in his ''
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genus, genera. It is the first work to consistently apply binomial nomenclature ...
'' of 1753. Two ''Equisetum'' plants are sold commercially under the names ''Equisetum japonicum'' (barred horsetail) and ''Equisetum camtschatcense'' (Kamchatka horsetail). These are both types of ''E. hyemale'', although they may also be listed as varieties of ''E. hyemale''.


Uses


Domestic

Boiled and dried ''Equisetum hyemale'' is used as traditional polishing material, similar to a fine grit
sandpaper upright=1.35, Sheets of sandpaper with different grit sizes (40 (coarse), 80, 150, 240, 600 (fine)) Sandpaper, also known as coated abrasive or emery paper, is a type of material that consists of sheets of paper or cloth with an abrasive substa ...
, in Japan. ;Music The stems are used to shape the reeds of reed instruments such as clarinets or saxophones.


Cultivation

''Equisetum hyemale'' cultivated as an
ornamental plant Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...
, for use in contained garden beds and planters, and in pots. It is a popular "icon plant" in contemporary Modernist and East Asian style
garden design Garden design is the art and process of designing and creating plans for layout and planting of garden, gardens and landscapes. Garden design may be done by the garden owner themselves, or by professionals of varying levels of experience and expe ...
. Its tight verticality fits into narrow planting spaces between walkways and walls, and on small balconies. It is also used as an accent plant in
garden pond A garden pond is a water feature constructed in a water garden or designed landscape, normally for aesthetic purposes, to provide wildlife habitat, or for swimming. Habitat Garden ponds can be excellent wildlife habitats and may contribute to ...
s and ornamental pools, and other landscape
water feature In landscape architecture and garden design, a water feature is one or more items from a range of fountains, jeux d'eau, pools, ponds, rills, artificial waterfalls, and streams. Before the 18th century they were usually powered by gravity ...
s, planted in submerged pots. The plant is sometimes sold in the nursery trade as "barred horsetail" or "''Equisetum japonicum''", but is different in appearance to '' Equisetum ramosissimum'' var. ''japonicum''.


Invasive behaviour

In
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
and
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
, the plant is an
invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
of moist natural
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s.


References

{{Authority control hyemale Plants described in 1753 Taxa named by Carl Linnaeus Ferns of Asia Ferns of Europe