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''Eocursor'' (meaning "dawn runner") is
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of basal
ornithischia Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
n
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
that lived in what is now
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
during the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
. Remains of this animal have been found in the
Upper Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, south ...
and it is among the most completely known early ornithischians, shedding new light on the origin of the group. The exact age of this taxon has been the subject of uncertainty. It was originally interprereted as living during the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
(
Norian The Norian is a division of the Triassic geological period, Period. It has the rank of an age (geology), age (geochronology) or stage (stratigraphy), stage (chronostratigraphy). It lasted from ~227.3 to Mya (unit), million years ago. It was prec ...
age), around 210
million years ago Million years ago, abbreviated as Mya, Myr (megayear) or Ma (megaannum), is a unit of time equal to (i.e. years), or approximately 31.6 teraseconds. Usage Myr is in common use in fields such as Earth science and cosmology. Myr is also used w ...
; however, Olsen, Kent & Whiteside (2010) stated that there is no independent geochronological support for its assumed age, and the available data makes it impossible to conclusively determine whether ''Eocursor'' is of Triassic or
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
(potentially as young as
Sinemurian In the geologic timescale, the Sinemurian is an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic epoch (geology), Epoch or series (stratigraphy), Series. It spans the time between 199.5 ±0.3 annu ...
) age. ''Eocursor'' was subsequently interpreted as a taxon of Early Jurassic age by McPhee ''et al.'' (2017), who identified the specimen as having originated from the Upper (not Lower) Elliot Formation. Fossils of ''Eocursor'' were originally collected in 1993, but were not formally described until fourteen years later. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, ''Eocursor parvus'', was described in 2007 by Richard J. Butler, Roger M. H. Smith, and David B. Norman. ''Eocursor'' was one of the earliest known ornithischians, and sheds some light on early dinosaur relationships because early dinosaurs are known from mostly incomplete skeletons. ''Eocursor'' is known from partial skeletal elements, including skull fragments, spinal elements, pelvis, long leg bones, and unusually large grasping hands.


Description

''Eocursor'' was a lightly built bipedal dinosaur with an estimated length of about 1 m (3 ft). The general aspect of the animal resembles that of the early Jurassic ornithischians such as ''
Lesothosaurus ''Lesothosaurus'' is a Monotypic taxon, monospecific genus of ornithischian dinosaur that lived during the Early Jurassic in what is now South Africa and Lesotho. It was named by paleontologist Peter Galton in 1978, the name meaning "lizard from L ...
'' and '' Scutellosaurus''. Its large hands resembled those of the
Heterodontosauridae Heterodontosauridae is a family (biology), family of ornithischian dinosaurs that were likely among the most Basal (phylogenetics), basal (primitive) members of the group. Their phylogenetic placement is uncertain but they are most commonly fou ...
, a
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
of primitive ornithischians. The morphology of the triangular teeth, not unlike those of an
iguana ''Iguana'' (, ) is a genus of herbivorous lizards that are native to tropical areas of Mexico, Central America, South America, and the Caribbean. The genus was first described by Austrian naturalist Josephus Nicolaus Laurenti, J.N. Laurenti in ...
, suggests partial herbivory. The
tibia The tibia (; : tibiae or tibias), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two Leg bones, bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outsi ...
was significantly longer than the
femur The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg. The Femo ...
, indicating it was a swift runner.


Discovery and naming

In 1993, the
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''Eocursor'' ( SAM-PK-K8025) was collected from the Upper Elliot Formation in Free State, South Africa. The skeleton, although only partially complete, is one of the best-preserved early ornithischian skeletons ever found. The fossils include a partial skeleton: portions of the skull, lower jaw, vertebrae, and limbs. In June 2007, the fossils were formally described by an international team of paleontologists: Richard Butler of the
Natural History Museum A natural history museum or museum of natural history is a scientific institution with natural history scientific collection, collections that include current and historical records of animals, plants, Fungus, fungi, ecosystems, geology, paleo ...
, London, and the
University of Cambridge The University of Cambridge is a Public university, public collegiate university, collegiate research university in Cambridge, England. Founded in 1209, the University of Cambridge is the List of oldest universities in continuous operation, wo ...
, David Norman of the
University of Cambridge The University of Cambridge is a Public university, public collegiate university, collegiate research university in Cambridge, England. Founded in 1209, the University of Cambridge is the List of oldest universities in continuous operation, wo ...
, and Roger M. H. Smith of the Iziko South African Museum. The fossils were named ''Eocursor parvus'', from the
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
word ''eos'' (meaning "dawn"), and the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
words ''cursor'' (meaning "runner") and ''parvus'' ("little"), "in reference to the early occurrence of this ornithischian, its apparent locomotory abilities and its small size."


Classification

''Eocursor'' was an early ornithischian, one of the first so-called "bird-hipped" dinosaurs, a group which would eventually give rise to animals such as ''
Stegosaurus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been fo ...
'', ''
Triceratops ''Triceratops'' ( ; ) is a genus of Chasmosaurinae, chasmosaurine Ceratopsia, ceratopsian dinosaur that lived during the late Maastrichtian age of the Late Cretaceous Period (geology), period, about 68 to 66 million years ago on the island ...
'', and ''
Iguanodon ''Iguanodon'' ( ; meaning 'iguana-tooth'), named in 1825, is a genus of iguanodontian dinosaur. While many species found worldwide have been classified in the genus ''Iguanodon'', dating from the Late Jurassic to Early Cretaceous, Taxonomy (bi ...
''. Butler ''et al.'' consider ''Eocursor'' more primitive than ''Lesothosaurus'' and Heterodontosauridae, but more derived than the ''
Pisanosaurus ''Pisanosaurus'' ( ) is an extinct genus of early dinosauriform, likely an ornithischian or silesaurid, from the Late Triassic of Argentina. It was a small, lightly built, ground-dwelling herbivore, that could grow up to an estimated long. On ...
'', as a basal ornithischian forming a sister
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
of
Genasauria Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek st ...
.


References


External links


''Eocursor parvus'' BBC article
{{Taxonbar , from=Q133498 Ornithischia Dinosaur genera Hettangian dinosaurs Elliot Formation Fossil taxa described in 2007 Taxa named by David B. Norman Dinosaurs of South Africa