The Entognatha, from

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

ἐντός (''entós''), meaning "inside", and γνάθος (''gnáthos''), meaning "jaw", are a

class Class, Classes, or The Class may refer to: Common uses not otherwise categorized * Class (biology), a taxonomic rank * Class (knowledge representation), a collection of individuals or objects * Class (philosophy), an analytical concept used d ...

of wingless and ametabolous

arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...

s, which, together with the

insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...

s, makes up the subphylum

Hexapoda The subphylum Hexapoda (from Greek for 'six legs') or hexapods comprises the largest clade of arthropods and includes most of the extant arthropod species. It includes the crown group class Insecta (true insects), as well as the much smaller c ...

. Their mouthparts are entognathous, meaning that they are retracted within the head, unlike the insects. Entognatha are apterous, meaning that they lack wings. The class contains three

orders Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * H ...

: Collembola (

springtail Springtails (class Collembola) form the largest of the three lineages of modern Hexapoda, hexapods that are no longer considered insects. Although the three lineages are sometimes grouped together in a class called Entognatha because they have in ...

s, 9000 species),

Diplura The order Diplura ("two-pronged bristletails") is one of three orders of non-insect hexapods within the class Entognatha (alongside Collembola (springtails) and Protura). The name "diplura", or "two tails", refers to the characteristic pair o ...

("two-tail", 1000 species) and

Protura The Protura, or proturans, and sometimes nicknamed coneheads, are very small (0.6–1.5mm long), soil-dwelling animals, so inconspicuous they were not noticed until the 20th century. The Protura constitute an order of hexapods that were previou ...

("first-tail", 800 species). These three groups were historically united with the now-obsolete order Thysanura to form the class

Apterygota The name Apterygota is sometimes applied to a former Subclass (biology), subclass of small, agile insects, distinguished from other insects by their lack of wings in the present and in their evolutionary history; notable examples are the silverfis ...

, but it has since been recognized that the hexapodous condition of these animals has evolved independently from that of insects, and independently ''within'' each order. The orders might not be closely related, and Entognatha is now considered to be a

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

group.

Morphology

These minute

s are apterous, unlike some orders of insects that have lost their wings secondarily (but are derived from winged ancestors). Their mouthparts are enclosed within a pouch in the head capsule, called the gnathal pouch, so only the tips of the mandibles and maxillae are exposed beyond the cavity. This pouch is created in the embryo by a flap or lateral head sclerite near the mouth on each side of the head which fuses with the labium. Other differences from insects are that each antennal segment is musculated; in insects, only the two basal segments are. Sperm transfer is always indirect, and there is an

ovipositor The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typica ...

in the females. Of the three orders, only collembolans possess eyes; nevertheless, many collembolans are blind, and even when

compound eye A compound eye is a Eye, visual organ found in arthropods such as insects and crustaceans. It may consist of thousands of ommatidium, ommatidia, which are tiny independent photoreception units that consist of a cornea, lens (anatomy), lens, and p ...

s are present, there are no more than eight ommatidia. *Collembolans have a ventral tube termed a collophore on the first abdominal segment. The collophore is involved in moisture absorption. On the third abdominal segment is the retinaculum that holds the furcula. The furcula is the "spring" for which the Collembola are given the name springtails. *Proturans, sometimes referred to as "coneheads", do not have eyes or antennae. They possess a

telson The telson () is the hindmost division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment (biology), segment of the arthropod body, or an additional division that is not a true segm ...

and abdominal styli thought to be vestigial legs. *Diplurans have a pair of caudal cerci, from which their name, meaning "two-tailed", is derived. They also possess abdominal styli.

References

{{Taxonbar, from=Q623286 Arthropod classes Paraphyletic groups