''Luciobarbus graellsii'' is a

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest verte ...

. It is here placed in ''

Luciobarbus ''Luciobarbus'' is a genus of ray-finned fishes in the family Cyprinidae. Its members are found in fresh and brackish waters of southern Europe, northern Africa, the wider Near East, the Aral and Caspian Seas, and rivers associated with these. Se ...

'' following the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

, but that

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

is very closely related to the other typical barbels and perhaps better considered a mere

subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...

of '' Barbus''. The Andalusian barbel was formerly included in '' L. bocagei'' as

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

.de Graaf ''et al.'' (2007), Almodóvar ''et al.'' (2008), Freyhof & Kottelat (2008) In addition, ''L. graellsii'' has – like its close relative the "Albanian barbel" (''

Luciobarbus albanicus The Albanian barbel (''Luciobarbus albanicus'') is a ray-finned fish species in the family Cyprinidae. It is often referred to as "Albanian barbel" in a literal translation of its scientific name. Distribution and ecology It occurs from the Mo ...

'') and the slightly more distantly related red-tailed barbel (''

Barbus haasi ''Barbus haasi'', or the "Catalonian barbel" ( ca, barb cua-roig; es, barbo colirrojo or ''barbo de cola roja''), is a species of freshwater fish in the family Cyprinidae. It is a small size barbel found only in the northeast of the Iberian pe ...

'') – also been placed in ''

Messinobarbus ''Barbus carottae'' is a species of ray-finned fish in the genus ''Barbus'' from Lake Yliki in Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of t ...

''. But even if that genus is valid, it is probably incorrect to do so.Freyhof & Kottelat (2008) It was originally

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to the northeast of

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...

, occurring chiefly on the

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...

side in the

Ebro , name_etymology = , image = Zaragoza shel.JPG , image_size = , image_caption = The Ebro River in Zaragoza , map = SpainEbroBasin.png , map_size = , map_caption = The Ebro ...

's to Ter River'

drainage basin A drainage basin is an area of land where all flowing surface water converges to a single point, such as a river mouth, or flows into another body of water, such as a lake or ocean. A basin is separated from adjacent basins by a perimeter, t ...

s, but also on the

Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...

side to the Asón drainage basin. Around 1998, it was introduced to some rivers of

Tuscany Tuscany ( ; it, Toscana ) is a Regions of Italy, region in central Italy with an area of about and a population of about 3.8 million inhabitants. The regional capital is Florence (''Firenze''). Tuscany is known for its landscapes, history, art ...

(

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

) and is now well established in the Albegna, Fiora and

Ombrone The Ombrone (Latin: ''Umbro'') is a long river in Tuscany, central Italy. The Ombrone's source is at San Gusmè, near Castelnuovo Berardenga, on the south-eastern side of the Monti del Chianti. After a twisting route, it receives the waters of t ...

drainage basins. ''L. graellsii'' is an

omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutr ...

, eating mainly large aquatic

invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...

s and

algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...

. They

spawn Spawn or spawning may refer to: * Spawn (biology), the eggs and sperm of aquatic animals Arts, entertainment, and media * Spawn (character), a fictional character in the comic series of the same name and in the associated franchise ** '' Spawn: Ar ...

from late spring to the height of summer (May to August). They

migrate Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...

upstream to their spawning sites, which are in faster and more shallow stretches of river, where the bottom is made of

gravel Gravel is a loose aggregation of rock fragments. Gravel occurs naturally throughout the world as a result of sedimentary and erosive geologic processes; it is also produced in large quantities commercially as crushed stone. Gravel is classifi ...

and rocks. This species becomes sexually mature at 4 years of age, with a

standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...

of . It is long-lived, and can get up to 16 years old. Abundant in its range, the Ebro Barbel not considered a

threatened species Threatened species are any species (including animals, plants and fungi) which are vulnerable to endangerment in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensa ...

by the

References

* (2008): Natural hybridization of ''Barbus bocagei'' x ''Barbus comizo'' (Cyprinidae) in Tagus River basin, central Spain nglish with French abstract ''Cybium'' 32(2): 99–102
PDF fulltext
* (2007): Evolutionary origin of Lake Tana's (Ethiopia) small ''Barbus'' species: indications of rapid ecological divergence and speciation. '' Anim. Biol.'' 57(1): 39–48. (HTML abstract) *

External links

Video of Ebro Barbels ''Luciobarbus graellsii'' on Youtube
graellsii Cyprinid fish of Europe Fish described in 1866 Taxa named by Franz Steindachner Taxonomy articles created by Polbot {{Cyprininae-stub