Domestic Yak

The yak (''Bos grunniens''), also known as the Tartary ox, grunting ox, hairy cattle, or domestic yak, is a species of long-haired domesticated cattle found throughout the Himalayan region, the Tibetan Plateau, Tajikistan, the Pamir Mountains, and as far north as Mongolia and Siberia. It is descended from the wild yak (''Bos mutus'').

Etymology

The English word ''yak'' originates from the . In Tibetan and Balti it refers only to the male of the species, the female being called or in Tibetan and in Balti. In English, as in most other languages that have borrowed the word, ''yak'' is usually used for both sexes, with ''bull'' or ''cow'' referring to each sex separately.

Taxonomy

Belonging to the genus '' Bos'', yaks are related to cattle (''Bos primigenius''). Mitochondrial DNA analyses to determine the evolutionary history of yaks have been inconclusive.

The yak may have diverged from cattle at any point between one and five million years ago, and there is some suggestion that it may be more closely related to bison than to the other members of its designated genus. Apparent close fossil relatives of the yak, such as '' Bos baikalensis'', have been found in eastern Russia, suggesting a possible route by which yak-like ancestors of the modern American bison could have entered the Americas.

The species was originally designated as ''Bos grunniens'' ("grunting ox") by Linnaeus in 1766. Still, this name is now generally considered to refer only to the domesticated form of the animal, with ''Bos mutus'' ("mute ox") being the preferred name for the wild species. Although some authors still consider the wild yak to be a subspecies, ''Bos grunniens mutus'', the ICZN made an official ruling in 2003 permitting the use of the name ''Bos mutus'' for wild yaks, and this is now the more common usage.

There are no recognised subspecies of yak except where the wild yak is considered a subspecies of ''Bos grunniens''.

Physical characteristics

Yaks are heavily built animals with bulky frames, sturdy legs, rounded, cloven hooves, and extremely dense, long fur hanging lower than the belly. While wild yaks are generally dark, blackish to brown in colouration, domestic yaks can be quite variable, often having rusty brown and cream patches. They have small ears and broad foreheads, with smooth horns that are generally dark in colour. In males (bulls), the horns sweep out from the sides of the head and then curve backwards; they typically range from in length.

The horns of females (cows) are smaller, at in length, and have a more upright shape. Both sexes have a short neck with a pronounced hump over the shoulders, although this is larger and more visible in males. Males weigh , females weigh . Wild (feral) yaks can be substantially heavier, bulls reaching weights of up to . Depending on the breed, domestic yak males are high at the withers, while females are high at the withers.

Both sexes have long, shaggy hair with a dense woolly undercoat over the chest, flanks, and thighs to insulate them from the cold. Especially in bulls, this may form a long "skirt" that can reach the ground. The tail is long and horselike rather than tufted like the tails of cattle or bison. Domesticated yaks have a wide range of coat colours, with some individuals being white, grey, brown, roan or piebald. The udder in females and the scrotum in males are small and hairy as protection against the cold. Females have four teats.

Yaks are not known to produce the characteristic lowing (mooing) sound of cattle, but both wild and domestic yaks grunt and squeak, which inspired the scientific name of the domestic yak variant, ''Bos grunniens'' (grunting bull). Nikolay Przhevalsky named the wild variant ''Bos mutus'' (silent bull), believing that it did not make a sound at all, but it does.

Physiology

Yak physiology is well adapted to high altitudes, having larger lungs and heart than cattle found at lower altitudes, as well as greater capacity for transporting oxygen through their blood, due to the persistence of foetal haemoglobin throughout life. Conversely, yaks have trouble thriving at lower altitudes, and are prone to suffering from heat exhaustion above about . Further adaptations to the cold include a thick layer of subcutaneous fat and an almost complete lack of functional sweat glands.

Compared with domestic cattle, the rumen of yaks is unusually large, relative to the omasum. This likely allows them to consume greater quantities of low-quality food at a time, and to ferment it longer to extract more nutrients. Yak consume the equivalent of 1% of their body weight daily while cattle require 3% to maintain condition. They are grazing herbivores, with their wild ancestors feeding primarily on grass and sedges, with some herbs and dwarf shrubs.

Reproduction and life history

Yaks mate in the summer, typically between July and September, depending on the local environment. For the remainder of the year, many bulls wander in small bachelor groups away from the large herds. Still, as the rut approaches, they become aggressive and regularly fight with each other to establish dominance. In addition to non-violent threat displays, bellowing, and scraping the ground with their horns, bull yaks compete more directly, repeatedly charging at each other with heads lowered or sparring with their horns. Like bison, but unlike cattle, males wallow in dry soil during the rut, often while scent-marking with urine or dung. Females enter oestrus up to four times a year, and females are receptive only for a few hours in each cycle.

Gestation lasts between 257 and 270 days, so that the young are born between May and June, and results in the birth of a single calf. The cow finds a secluded spot to give birth, but the calf can walk within about ten minutes of birth, and the pair soon rejoin the herd. Females of both the wild and domestic forms typically give birth only once every other year, although more frequent births are possible if the food supply is good.

Calves are weaned at one year and become independent shortly thereafter. Wild calves are initially brown in color and only later develop darker adult hair. Females generally give birth for the first time at three or four years of age, and reach their peak reproductive fitness at around six years. Yaks may live for more than twenty years in domestication or captivity, although it is likely that this may be somewhat shorter in the wild.

Husbandry

For thousands of years, domesticated yaks have been kept in Mongolia and Tibet, primarily for their milk, fibre (wool), and meat, and as beasts of burden. Their dried droppings are an important fuel, used all over Tibet, and are often the only fuel available on the high, treeless Tibetan Plateau. Yaks transport goods across mountain passes for local farmers and traders and are an attraction for climbing and trekking expeditions: "Only one thing makes it hard to use yaks for long journeys in barren regions. They will not eat grain, which could be carried on the journey. They will starve unless they can be brought to a place where there is grass." They also are used to draw ploughs. Yaks' milk is often processed to a cheese called '' chhurpi'' in Tibetan and Nepali languages, and '' byaslag'' in Mongolia. Butter made from yaks' milk is an ingredient of the butter tea that Tibetans consume in large quantities, and is also used in lamps and made into butter sculptures used in religious festivities.

Outside the Himalayas

Small numbers of herds can be found in the United States, Canada, New Zealand, and some parts of Europe. Yaks have generated interest outside the Himalayas as a commercial crop and by cattle breeders. The main interest of North American yak breeders is lean meat production by hybridizing with other cattle, followed by wool production.

Research

The Indian government established a dedicated centre for research into yak husbandry, the ICAR-National Research Centre on Yak, in 1989. It is located at Dirang, Arunachal Pradesh, and maintains a yak farm in the Nyukmadung area at an altitude of above MSL.

Breeding and hybridization

In Nepal, Tibet, and Mongolia, domestic cattle are crossbred with yaks. This gives rise to the infertile male dzo མཛོ། as well as fertile females known as or མཛོ་མོ།, which may be crossed again with cattle. The Dwarf Lulu breed, "the only ''Bos primigenius taurus'' type of cattle in Nepal" has been tested for DNA markers and found to be a mixture of both taurine and zebu types of cattle (''B. p. taurus'' and ''B. p. indicus'') with yak. According to the International Veterinary Information Service, the low productivity of second-generation cattle–yak crosses makes them suitable only as meat animals.

Crosses between yaks and domestic cattle (''Bos primigenius taurus'') have been recorded in Chinese literature for at least 2,000 years. Successful crosses have also been recorded between yak and American bison,Zhang, R.C. (14 December 2000)
"Interspecies Hybridization between Yak, ''Bos taurus'' and ''Bos indicus'' and Reproduction of the Hybrids"
In: ''Recent Advances in Yak Reproduction'', Zhao, X.X.; Zhang, R.C. (eds.). International Veterinary Information Service. gaur, and banteng, generally with similar results to those produced with domestic cattle.

Domestication

Jacques et al. (2021) show that most elaborate yak-related terminologies are found within Tibetic and Gyalrongic languages. Both branches also have native terms for yak-cattle hybrids, suggesting that Tibetic and Gyalrongic speakers may have independently cross-bred yaks and cattle, predating the proto-Gyalrongic split (3221 169–4319 BP) from Tibeto-Gyalrongic. The oldest dated physical evidence of yak domestication is from 2,500 years BP.

Customs

In parts of Tibet and Karakorum, yak racing is a form of entertainment at traditional festivals and an important part of their culture. More recently, sports involving domesticated yaks, such as yak skiing or yak polo, are being marketed as tourist attractions in South Asian countries, including in Gilgit-Baltistan, Pakistan.

In Nepal, an annual festival is held to drink the fresh blood of yak, and it is believed that it cures various diseases such as gastritis, jaundice, and body strain. The fresh blood is extracted from the neck of a yak without killing it. The cut is healed after the ceremony is over. The ritual is believed to be originated in Tibet and Mustang.

Traditionally, yaks are used to transport deceased people to charnel grounds for sky burials in Tibet, with Tibetan practice holding that the yaks be set free upon the conclusion of the task.

Gallery

File:Yaks in Manali.jpg, Yaks in Manali, Himachal Pradesh, India, saddled for riding
File:LitangMonastery.jpg, Train of pack yaks at Litang monastery in Garzê Tibetan Autonomous Prefecture, Sichuan, China
File:Yaks still provide the best way to plow fields in Tibet.jpg, Yaks plowing fields in Tibet
File:YakRace.jpg, Yaks in Gilgit-Baltistan, Pakistan
File:四川 阿坝 白牦牛 - panoramio.jpg, Domestic yak in Mao County, China
File:四姑娘山景区 Mount Siguniang Scenic Area 17.jpg, A yak at Mount Siguniang Scenic Area, Sichuan, China

See also

* Yakalo
* Yak dance

References

External links

*
*
International Yak Association (IYAK)European Yak Association (EYAK)Yak Genome Database

{{Taxonbar, from=Q19829410

Livestock
Domesticated animals
Pack animals
Mammals of Central Asia
Mammals of East Asia
Mammals of Siberia
Mammals of South Asia
Mammals of Tibet
Fauna of the Himalayas
Mammals described in 1766
Taxa named by Carl Linnaeus
Bos