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''Cunninghamia'' is a genus of one or two living species of
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
coniferous Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s in the cypress family
Cupressaceae Cupressaceae or the cypress family is a family of conifers. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioecious or (rarely) dioecio ...
. They are native to China, northern Vietnam and Laos, and perhaps also Cambodia. They may reach in height. In vernacular use, it is most often known as ''Cunninghamia'', but is also sometimes called "China-fir" (though it is not a
fir Firs are evergreen coniferous trees belonging to the genus ''Abies'' () in the family Pinaceae. There are approximately 48–65 extant species, found on mountains throughout much of North and Central America, Eurasia, and North Africa. The genu ...
). The genus name ''Cunninghamia'' honours Dr. James Cunningham, a
British British may refer to: Peoples, culture, and language * British people, nationals or natives of the United Kingdom, British Overseas Territories and Crown Dependencies. * British national identity, the characteristics of British people and culture ...
doctor who introduced this species into cultivation in 1702 and botanist Allan Cunningham. A female cone Cluster of male cones


Description

The general shape of the tree is conical with tiered, horizontal
branch A branch, also called a ramus in botany, is a stem that grows off from another stem, or when structures like veins in leaves are divided into smaller veins. History and etymology In Old English, there are numerous words for branch, includ ...
es that are often somewhat pendulous toward the tips. ''Cunninghamia'' bears softly spined, leathery, stiff, green to blue-green needle-like
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
that spiral around the
stem Stem or STEM most commonly refers to: * Plant stem, a structural axis of a vascular plant * Stem group * Science, technology, engineering, and mathematics Stem or STEM can also refer to: Language and writing * Word stem, part of a word respon ...
with an upward arch; they are 2–7cm long and 3–5mm broad at the base, and bear two white or greenish white
stomata In botany, a stoma (: stomata, from Greek ''στόμα'', "mouth"), also called a stomate (: stomates), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange between the internal air spa ...
l bands underneath and sometimes also above. The foliage may turn bronze-tinted in very cold winter weather. The
cones In geometry, a cone is a three-dimensional figure that tapers smoothly from a flat base (typically a circle) to a point not contained in the base, called the ''apex'' or '' vertex''. A cone is formed by a set of line segments, half-lines, ...
are small and inconspicuous at
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds. Pollinating agents can be animals such as insects, for example bees, beetles or bu ...
in late winter, the pollen cones in clusters of 10–30 together, the female cones singly or 2–3 together. The seed cones mature in 7–8 months to 2.5–4.5cm long, ovoid to globose, with spirally arranged scales; each scale bears 3–5
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s. They are often proliferous (with a vegetative shoot growing on beyond the tip of the cone) on cultivated trees; this is rare in wild trees, and may be a
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
selected for easy vegetative propagation for use in
forestry Forestry is the science and craft of creating, managing, planting, using, conserving and repairing forests and woodlands for associated resources for human and Natural environment, environmental benefits. Forestry is practiced in plantations and ...
plantation Plantations are farms specializing in cash crops, usually mainly planting a single crop, with perhaps ancillary areas for vegetables for eating and so on. Plantations, centered on a plantation house, grow crops including cotton, cannabis, tob ...
s. As the tree grows its trunk tends to sucker around the base, particularly following damage to the stem or roots, and it then may grow in a multi-trunked form. Brown
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
of mature trees peels off in strips to reveal reddish-brown inner bark. Older specimens often look ragged, as the old needles may cling to stems for up to 5 years. Although it resembles conifer species found in much colder climates, the tree is highly tolerant of heat and can be found growing down to sea level in regions such as Hong Kong.


Taxonomy

It is the most basal extant member of the Cupressaceae. Phylogenetic evidence indicates that its lineage diverged from the rest of the family during the early
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
. Fossil representatives of the subfamily Cunninghamioideae, of which ''Cunninghamia'' is the only living member, extend back to the Middle Jurassic. The subfamily, which has 12 described genera, experienced a high diversity during the Cretaceous, but underwent a severe decline at the end of the Cretaceous, leaving ''Cunninghamia'' as the only living genus by the beginning of the Cenozoic. In the past, the genus was usually treated in the family Taxodiaceae, but this family is now included within the Cupressaceae. A few botanists have also treated it in a family of its own, Cunninghamiaceae, but this is not widely followed. The oldest fossil species of the genus are from the Late Cretaceous (
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
) of North America, including ''Cunninghamia hornbyensis'' from British Columbia, Canada, and ''Cunninghamia taylorii'' from the Horseshoe Canyon Formation of Alberta, Canada. Other fossils are known from the
Cenozoic The Cenozoic Era ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three g ...
of Asia, North America and Europe. The genus is traditionally said to contain two similar species, ''Cunninghamia lanceolata'' and ''C. konishii'', often referred to as the China fir and Taiwan fir, respectively. ''C. lanceolata'' occurs in mainland
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
,
Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
, and
Laos Laos, officially the Lao People's Democratic Republic (LPDR), is the only landlocked country in Southeast Asia. It is bordered by Myanmar and China to the northwest, Vietnam to the east, Cambodia to the southeast, and Thailand to the west and ...
, whereas ''C. konishii'' is restricted to
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...
. However, molecular genetic evidence is suggesting that they are the same species, and that ''C. konishii'' of Taiwan derive from multiple colonizations from the mainland. As ''C. lanceolata'' was the first name published, this name takes priority if the two are combined. In that case, Taiwan fir becomes ''Cunninghamia lanceolata'' var. ''konishii''. However, there is no consensus yet as to whether the two species should be combined.


Usage

''Cunninghamia'' is a prized
timber Lumber is wood that has been processed into uniform and useful sizes (dimensional lumber), including beams and planks or boards. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, window frames). ...
tree in China, producing soft, highly durable scented
wood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
similar to that of Coast Redwood and
Sugi ''Cryptomeria'' (literally "hidden parts") is a monotypic genus of conifer in the cypress family Cupressaceae. It includes only one species, ''Cryptomeria japonica'' ( syn. ''Cupressus japonica'' L.f.). It is considered to be endemic to Japan ...
. It is used in particular for manufacture of
coffin A coffin or casket is a funerary box used for viewing or keeping a corpse, for burial, entombment or cremation. Coffins are sometimes referred to as caskets, particularly in American English. A distinction is commonly drawn between "coffins" a ...
s and in
temple A temple (from the Latin ) is a place of worship, a building used for spiritual rituals and activities such as prayer and sacrifice. By convention, the specially built places of worship of some religions are commonly called "temples" in Engli ...
building where the scent is valued. ''Cunninghamia'' is grown as an ornamental tree in
park A park is an area of natural, semi-natural or planted space set aside for human enjoyment and recreation or for the protection of wildlife or natural habitats. Urban parks are urban green space, green spaces set aside for recreation inside t ...
s and large
garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate bot ...
s, where it typically reaches a height of 15–30 m. Due to its heat tolerance, it has been used as a
Christmas tree A Christmas tree is a decorated tree, usually an evergreen pinophyta, conifer, such as a spruce, pine or fir, associated with the celebration of Christmas. It may also consist of an artificial tree of similar appearance. The custom was deve ...
in subtropical areas such as the southern USA.


References


External links


Arboretum de Villardebelle: photo of cone

Arboretum de Villardebelle: photo of tree
{{Authority control Cupressaceae Conifer genera