Crichtonpelta
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''Crichtonpelta'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of extinct herbivorous
ankylosaurid Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pal ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
) of
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
.


Discovery and naming

In 2007,
Lü Junchang Lü Junchang (; 1965 – 9 October 2018) was a Chinese palaeontologist and professor at the Institute of Geology, Chinese Academy of Geological Sciences. An expert on Mesozoic reptiles, he described and named dozens of dinosaur and pterosaur taxa ...
, Ji Qiang, Gao Yubo and Li Zhixin named and described a second species of '' Crichtonsaurus'': ''Crichtonsaurus benxiensis''. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
refers to the Benxi Geological Museum. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
, BXGMV0012, is a skull found near
Beipiao Beipiao () is a city in Chaoyang prefecture, Liaoning province, in Northeast China. It has a population of 202,807. The main industry in the area is coal mining. With vertical shafts of almost 1000m, these are some of the deepest coal mines in C ...
in a layer of the Sunjiawan Formation probably dating from the early Cenomanian (~99–95 Ma). Specimen BXGMV0012-1, a skeleton lacking the skull, discovered in the same quarry as the holotype, was referred to the species. Furthermore, a skeleton with skull, displayed by the Sihetun Fossil Museum as a ''Crichtonsaurus bohlini'' specimen, was in 2014 referred to ''Crichtonpelta''. In 2017, a fourth specimen was described, from the same quarry as the holotype, G20090034, consisting of a skull lacking the front snout.YANG Jingtao, YOU Hailu, XIE Li & ZHOU Hongrui, 2017, "A New Specimen of ''Crichtonpelta benxiensis'' (Dinosauria: Ankylosaurinae) from the Mid-Cretaceous of Liaoning Province, China", ''Acta Geologica Sinica'' 91(3): 781-790 In 2014, Victoria Arbour concluded that ''Crichtonsaurus'' were a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
''. Therefore, she named a separate genus for its second species: ''Crichtonpelta''. The generic name combines a reference to
Michael Crichton John Michael Crichton (; October 23, 1942 – November 4, 2008) was an American author, screenwriter and filmmaker. His books have sold over 200 million copies worldwide, and over a dozen have been adapted into films. His literary works heavil ...
, the author of ''
Jurassic Park ''Jurassic Park'', later referred to as ''Jurassic World'', is an American science fiction media franchise created by Michael Crichton, centered on a disastrous attempt to create a theme park of De-extinction#Cloning, cloned dinosaurs. It bega ...
'', with a Greek πέλτη, ''peltè'', "small shield". At the time this was an invalid ''nomen ex dissertatione''. However, in 2015, ''Crichtonpelta'' was validly named by Arbour and Philip John Currie. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Crichtonsaurus benxiensis''; the ''
combinatio nova In biological taxonomy, a combinatio nova (abbreviated comb. nov. or n. comb.) refers to the formal renaming of an organism's scientific name when it is transferred to a different genus, reclassified within a different species, or its taxonomic ...
'' is ''Crichtonpelta benxiensis''. There was a possibility that, though ''Crichtonsaurus bohlini'' was a ''nomen dubium'', its fossil material in fact belonged to ''Crichtonpelta''. Arbour however, noted clear differences in the scapula and humerus between BXGMV0012-1 and LPM 101, a specimen previously referred to ''Crichtonsaurus bohlini'': the scapula of the former has a tab-like
acromion In human anatomy, the acromion (from Greek: ''akros'', "highest", ''ōmos'', "shoulder", : acromia) or summit of the shoulder is a bony process on the scapula (shoulder blade). Together with the coracoid process, it extends laterally over the sh ...
and its humerus a much longer deltopectoral crest.


Description

Arbour established several distinguishing traits. One of these was an
autapomorphy In phylogenetics, an autapomorphy is a distinctive feature, known as a Synapomorphy, derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or Outgroup (cladistics), outgroup taxa, not ...
, unique derived trait: the apex of the (quadrato)jugal, or cheek, horn, is pointing upwards. Also a unique combination of in themselves not unique traits is present. The upper snout armour forms an amorphous mass, not clearly separated into distinctive tiles. The
jugal bone The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic bone, zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by spe ...
is deeper than that of ''
Pinacosaurus ''Pinacosaurus'' (meaning "Plank lizard") is a genus of ankylosaurid thyreophoran dinosaur that lived in Asia during the Late Cretaceous (Campanian, roughly 75 to 71 million years ago), mainly in Mongolia and China. The first remains of the genu ...
''. The skull roof is not notched at the
lacrimal bone The lacrimal bones are two small and fragile bones of the facial skeleton; they are roughly the size of the little fingernail and situated at the front part of the medial wall of the orbit. They each have two surfaces and four borders. Several bon ...
as in ''Pinacosaurus grangeri''. The squamosal horns are shorter than those of ''Pinacosaurus mephistocephalus''. However, these horns are longer and more pointed than those of ''
Gobisaurus ''Gobisaurus'' is an extinct genus of herbivorous basal ankylosaurid ankylosaur from the Lower Cretaceous of China (''Nei Mongol Zizhiqu''). The genus is monotypic, containing only the species ''Gobisaurus domoculus''. Discovery and naming Th ...
'' or ''
Shamosaurus ''Shamosaurus'' is an extinct genus of herbivorous basal ankylosaurid ankylosaur from Early Cretaceous (Aptian to Albian stage) deposits of Höövör, Mongolia. Discovery and naming In 1977, a Soviet-Mongolian expedition discovered the skelet ...
''. The point of the cheek horn is located on the rear edge. The transverse crest on the top of the rear skull has two points. The holotype of ''Crichtonpelta'' is somewhat larger than ''Crichtonsaurus'', itself about three to four metres long. It is uncertain whether ''Crichtonpelta'' already possessed a tail club.


Classification

''Crichtonpelta'' was, within the
Ankylosauridae Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pal ...
, placed in the Ankylosaurinae, in a basal position. If correct, this makes it the oldest known ankylosaurine.


References

{{Taxonbar, from=Q20737863 Ankylosaurinae Dinosaur genera Cenomanian dinosaurs Taxa named by Philip J. Currie Fossil taxa described in 2015 Dinosaurs of China