The family Corydalidae contains the
megalopterous insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s known as
dobsonflies
Dobsonflies are a subfamily of insects, Corydalinae, part of the Megalopteran family Corydalidae. The larvae (commonly called hellgrammites) are aquatic, living in streams, and the adults are often found along streams as well. The nine genera ...
and
fishflies. Making up about three dozen
genera
Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
, they occur primarily throughout
North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
, both temperate and tropical,
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
,
Australia
Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
,
New Zealand
New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
,
Africa
Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
(particularly
South Africa
South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
) and
Asia
Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
.
They are sizeable Megaloptera, with a body usually larger than 25 mm (1 inch). They often have long filamentous
antennae, though in male fishflies they are characteristically feathered.
Ocelli
A simple eye or ocellus (sometimes called a pigment pit) is a form of eye or an optical arrangement which has a single lens without the sort of elaborate retina that occurs in most vertebrates. These eyes are called "simple" to distinguish the ...
are present; the fourth
tarsal segment is
cylinder
A cylinder () has traditionally been a three-dimensional solid, one of the most basic of curvilinear geometric shapes. In elementary geometry, it is considered a prism with a circle as its base.
A cylinder may also be defined as an infinite ...
-shaped. The four large
wings are translucent, smoky grey, or mixed, and the
anterior
Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
pair is slightly longer than the posterior one. Their aquatic larvae are used as fish bait and are called hellgrammites.
The
larva
A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e are aquatic, active, armed with strong sharp mandibles, and breathe by means of abdominal
branchial filaments. When full sized — which can take several years — they leave the water and spend a quiescent
pupa
A pupa (; : pupae) is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages th ...
l stage on the land, in chambers dug under stones or logs, before
metamorphosis
Metamorphosis is a biological process by which an animal physically develops including birth transformation or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and different ...
into the sexually mature insect.
File:Parachauliodes japonicus 01 larva.JPG, ''Parachauliodes japonicus''
File:Corydalis larva Packard.jpg, Larva
Distribution
The eastern dobsonfly, ''
Corydalus cornutus'', is the most well-known
North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
n
species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
among the dobsonflies. These
genera
Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
have distinctive elongated mandibles in males and form the
subfamily
In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Corydalinae. The genera in which the males have normal mandibles, called fishflies, form the subfamily
Chauliodinae. The summer fishfly, ''
Chauliodes pectinicornis'', is perhaps the best-known of these in North America; its immense mating swarms in the
Upper Mississippi River
The Upper Mississippi River is the portion of the Mississippi River upstream of St. Louis, Missouri, United States, a city at the confluence of its main tributary, the Missouri River. Historically, it may refer to the area above the Arkansa ...
region fill the air on a few summer nights each year much like
mayflies
Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the order ...
in certain regions of
Europe
Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
, leaving millions of carcasses to be cleaned up the next day.
File:Chauliodes rastricornis P1400671a.jpg, '' Chauliodes pectinicornis''
File:CorydalisAsiaticaParkerCoward.jpg, '' Acanthacorydalis asiatica''
Taxonomy
The family is divided into two subfamilies, the Corydalinae and the Chauliodinae. Classification and identification is based on external morphology and at the species level using male genitalia.
These 36 genera belong to the family Corydalidae:
* ''
Acanthacorydalis''
c g
* ''
Anachauliodes''
c g
* ''
Apochauliodes''
c g
* ''
Archichauliodes''
c g
* ''
Chauliodes''
Latreille, 1796 i c g b – fishflies
* ''
Chauliosialis''
c g
* ''
Chloronia''
Banks, 1908 i c g
* ''
Chloroniella''
c g
* ''
Corydalites''
c g
* ''
Corydalus''
Latreille, 1802 i c g b – dobsonflies
* ''
Cratocorydalopsis''
c g
* ''
Cretochaulus''
c g
* ''
Ctenochauliodes''
c g
* ''
Dysmicohermes''
Munroe, 1953 i c g b
* ''
Eochauliodes''
c g
* ''
Jurochauliodes''
c g
* ''
Lithocorydalus''
c g
* ''
Madachauliodes''
c g
* ''
Neochauliodes''
c g
* ''
Neohermes''
Banks, 1908 i c g b
* ''
Neoneuromus''
c g
* ''
Neurhermes''
c g
* ''
Nevromus''
c g
* ''
Nigronia''
Banks, 1908 i c g b – dark fishflies
* ''
Nothochauliodes''
c g
* ''
Orohermes''
Evans, 1984 i c g b
* ''
Parachauliodes''
c g
* ''
Platychauliodes''
c g
* ''
Platyneuromus''
Weele, 1909 i c g
* ''
Protochauliodes''
Weele, 1909 i c g b
* ''
Protohermes
''Protohermes'' is a genus of Dobsonfly, dobsonflies in the family Corydalidae. ''Protohermes'' is the most speciose and widely distributed genus within Megaloptera, but up to 85% of species are restricted to small Endemism, endemic areas in Asi ...
''
van-der Weele, 1907 g
* ''
Puri
Puri, also known as Jagannath Puri, () is a coastal city and a Nagar Palika, municipality in the state of Odisha in eastern India. It is the district headquarters of Puri district and is situated on the Bay of Bengal, south of the state ca ...
''
c g
* ''
Taeniochauliodes''
c g
Data sources: i = ITIS,[ c = Catalogue of Life,][ g = GBIF,][ b = Bugguide.net]
Footnotes
References
External links
Informative Corydalidae video*
*
{{Authority control
Megaloptera
Insect families