Coniopterygidae
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The dustywings, Coniopterygidae, are a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
Pterygota Pterygota ( ) is a subclass of insects that includes all winged insects and groups who lost them secondarily. Pterygota group comprises 99.9% of all insects. The orders not included are the Archaeognatha (jumping bristletails) and the Zygent ...
(winged insects) of the net-winged insect
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
(
Neuroptera The insect order (biology), order Neuroptera, or net-winged insects, includes the lacewings, mantidflies, antlions, and their relatives. The order consists of some 6,000 species. Neuroptera is grouped together with the Megaloptera (alderflies, f ...
). About 460 living
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
are known.Engel & Grimaldi (2007) These tiny insects can usually be determined to
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
with a
hand lens A magnifying glass is a convex lens—usually mounted in a frame with a handle—that is used to produce a magnification, magnified image of an object. A magnifying glass can also be used to focus light, such as to concentrate the Sun's radiati ...
according to their wing venation, but to distinguish
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
, examination of the
genital A sex organ, also known as a reproductive organ, is a part of an organism that is involved in sexual reproduction. Sex organs constitute the primary sex characteristics of an organism. Sex organs are responsible for producing and transporting ...
s by
microscope A microscope () is a laboratory equipment, laboratory instrument used to examine objects that are too small to be seen by the naked eye. Microscopy is the science of investigating small objects and structures using a microscope. Microscopic ...
is usually necessary.


Description and ecology

In general habitus, the adults are quite unlike other net-winged insects. Because of their small size - wingspan is between 1.8 and 5 millimetres - and their translucent brownish wings usually covered with the namesake whitish dust of
wax Waxes are a diverse class of organic compounds that are lipophilic, malleable solids near ambient temperatures. They include higher alkanes and lipids, typically with melting points above about 40 °C (104 °F), melting to give lo ...
y scales, they may at first be mistaken for whiteflies (
Aleyrodidae Whiteflies are Hemipterans that typically feed on the undersides of plant leaves. They comprise the family (biology), family Aleyrodidae, the only family in the superfamily Aleyrodoidea. More than List of whitefly species, 1550 species have been ...
). But whiteflies are
true bug Hemiptera (; ) is an order of insects, commonly called true bugs, comprising more than 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from to a ...
s (Sternorrhyncha), which are only distantly related to net-winged insects. An easily perceived distinguishing feature to separate whiteflies from dustywings is, that like many other Neuroptera, dustywings carry their wings nearly side-by-side when at rest, whereas whiteflies carry them almost flat across the back. There are no more than two veins across the
costa Costa may refer to: Biology * Rib (Latin: ''costa''), in vertebrate anatomy * Costa (botany), the central strand of a plant leaf or thallus * Costa (coral), a stony rib, part of the skeleton of a coral * Costa (entomology), the leading edge of th ...
l field, and few cross-veins in general - unique among the living net-winged insects, dustywings do not actually have the "net-winged" venation. Some Coniopterygidae, like the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Conwentzia ''Conwentzia'' is a genus of net-winged insects belonging to the family Coniopterygidae or ''dustywings''. ''Conwentzia'' species are present in most of Europe. These tiny insects, also known as waxwing lacewings or dustywing lacewings. They hav ...
'', have only vestigial hindwings; others, like ''
Helicoconis ''Helicoconis'' is a genus of insects belonging to the family Coniopterygidae The dustywings, Coniopterygidae, are a family (biology), family of Pterygota (winged insects) of the net-winged insect order (biology), order (Neuroptera). About 460 ...
'' females, are completely wingless. Dustywings are strongly associated with woody
plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
s, on and around which they usually spend their entire lives. Females deposit their eggs singly on
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
or
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
. Dustywing
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e are around 3.5 mm long. Their mouthparts consist of short, straight sucking tubes covered by the labrum (upper "lip"). They are
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal (active during dawn), vespertine (biology), vespertine/vespertinal (active during dusk), or both. This is distinguished from diurnalit ...
and dwell on
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
s and
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s, where they feed on small
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s like
scale insect Scale insects are small insects of the Order (biology), order Hemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient g ...
s,
aphid Aphids are small sap-sucking insects in the Taxonomic rank, family Aphididae. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white Eriosomatinae, woolly ...
s and
mite Mites are small arachnids (eight-legged arthropods) of two large orders, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari. However, most recent genetic analyses do not recover the two as eac ...
s, as well as on
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
eggs; the mouth tubes are used for sucking fluids from the prey. There are usually two
generation A generation is all of the people born and living at about the same time, regarded collectively. It also is "the average period, generally considered to be about 20–⁠30 years, during which children are born and grow up, become adults, and b ...
s each year.


Systematics and taxonomy

Due to the dustywings' many
apomorph In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to ha ...
ies, the
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Coniopterygoidea was formerly believed to be
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
, and the primitive traits of their larvae were held to evidence a quite basal place among the net-winged insects. But in fact the spongillaflies (
Sisyridae Sisyridae, commonly known as spongeflies or spongillaflies, are a family of winged insects in the order Neuroptera. There are approximately 60 living species described, and several extinct species identified from the fossil record. Description ...
), formerly allied with the
Osmylidae Osmylidae are a small family of winged insects of the net-winged insect order Neuroptera. The osmylids, also called lance lacewings, stream lacewings or giant lacewings, are found all over the world except North and Central America. There are ar ...
in error due to their larvae's convergent
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
, seem to be close relatives of the Coniopterygidae, more
plesiomorph In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral Phenotypic trait, character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorph ...
ic altogether as adults but with a number of peculiar and highly divergent apomorphies, particularly in the larvae. So even though the spongillaflies are not generally placed in the Coniopterygoidea yet, they most likely form a
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
with the dustywings and thus it would seem that the Coniopterygoidea, rather than being maintained as an unnecessarily
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
, are better expanded to signify that the spongillaflies and the dustywings are each other's closest relatives among the net-winged insects. This is all the more significant because in this apparent clade, there would be a highly interesting and exactly opposing pattern of
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
- primitive larvae and highly advanced adults in the dustywings, versus primitive adults and very advanced larvae in the spongillaflies.See references in Haaramo (2008)


Genera


Living genera

* '' Aleuropteryx'' Löw, 1855 * '' Brucheiser'' Riek, 1975 * '' Coniocompsa'' Enderlein, 1905 * '' Coniopteryx'' Curtis, 1834 * ''
Conwentzia ''Conwentzia'' is a genus of net-winged insects belonging to the family Coniopterygidae or ''dustywings''. ''Conwentzia'' species are present in most of Europe. These tiny insects, also known as waxwing lacewings or dustywing lacewings. They hav ...
'' Enderlein, 1905 * '' Cryptoscenea'' Enderlein, 1914 * '' Flintoconis'' Sziráki, 2007 * ''
Helicoconis ''Helicoconis'' is a genus of insects belonging to the family Coniopterygidae The dustywings, Coniopterygidae, are a family (biology), family of Pterygota (winged insects) of the net-winged insect order (biology), order (Neuroptera). About 460 ...
'' Enderlein, 1905 * '' Hemisemidalis'' Meinander, 1972 * '' Heteroconis'' Enderlein, 1905 * '' Neoconis'' Enderlein, 1930 * '' Neosemidali''s Enderlein, 1930 * '' Nimboa'' Navas, 1915 * '' Parasemidalis'' Enderlein, 1905 * '' Semidalis'' Enderlein, 1905 * '' Spiloconis'' Enderlein, 1907 * '' Vartiana'' H. Aspock & U. Aspock, 1965


Extinct genera

The earliest coniopterygid is '' Toarciconiopteryx'' from the Early Jurassic (
Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...
) Green Series of Germany, dating to around 182 million years ago, with numerous other fossil species known, predominantly in amber. The supposed
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic ...
dustywing genus '' Archiconiopteryx'' actually seems to be a whitefly-like
true bug Hemiptera (; ) is an order of insects, commonly called true bugs, comprising more than 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from to a ...
. Subfamily Aleuropteryginae * †''Burmaleuropteryx'' Bai et al. 2019
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
, Myanmar, Late Cretaceous (
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
) * †''Garnaconis'' Perrichot and Nel 2014 Vendée amber, France, Late Cretaceous (
Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...
) * †''Juraconiopteryx'' Meinander 1975
Karabastau Formation The Karabastau Formation () is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that has been studied since ...
, Kazakhstan, Middle-Late Jurassic (
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma (million years ago) and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
/ Oxfordian) * †''Libanoconis'' Engel 2002
Lebanese amber Lebanese amber is fossilized resin found in Lebanon and its surroundings. It dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. It formed on what was then the northern coast of Gondwana, believed to be a tropic ...
, Early Cretaceous (
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
),
Taimyr amber Taymyr or Taimyr may refer to: Places *Taymyr Peninsula, a peninsula in Siberia * Taymyr Gulf *Taymyra, a river in the Taymyr Peninsula * Lake Taymyr * Taymyr Island, an island in the Kara Sea *Taymyr Autonomous Okrug, a former federal subject of ...
,
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
* †''Palaeoconis'' Ružičková et al. 2019 Burmese amber, Myanmar, Cenomanian * Tribe Coniocompsini Enderlein 1905 ** †''Archiconiocompsa'' Enderlein 1910
Baltic amber Baltic amber or succinite is amber from the Baltic region, home of its largest known deposits. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that this forested region provided the re ...
,
Rovno amber Rovno amber, occasionally called Ukrainian amber, is amber found in the Rivne Oblast and surrounding regions of Ukraine and Belarus. The amber is dated between Late Eocene and Early Oligocene, and suggested to be contemporaneous to Baltic amber ...
,
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
** †''Geroconiocompsa'' Engel 2010 Balic amber, Eocene * Tribe Fontenelleini ** †''Achlyoconis'' Engel 2016 Burmese amber, Myanmar, Cenomanian ** †''Alboconis'' Nel et al. 2005 Charentese amber, France, Cenomanian **†''Archiconis'' Enderlein 1930 Baltic amber, Eocene **†''Cycloconis'' Li et al. 2019 Burmese amber, Myanmar, Cenomanian **†''Glaesoconis'' Meinander 1975 Burmese amber, Myanmar, Cenomanian,
New Jersey amber New Jersey Amber, sometimes called Raritan amber, is amber found in the Raritan Formation, Raritan and Magothy Formations of the Mid-Atlantic states, Central Atlantic (Eastern) coast of the United States. It is dated to the Late Cretaceous, Turoni ...
,
Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...
Taimyr amber, Late Cretaceous (
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...
) **†''Pararchiconis'' Nel 1991 Alsace, France,
Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...
(
Rupelian The Rupelian is, in the geologic timescale, the older of two age (geology), ages or the lower of two stage (stratigraphy), stages of the Oligocene epoch (geology), Epoch/series (stratigraphy), Series. It spans the time between . It is preceded b ...
) **†''Soplaoconis'' Pérez-de la Fuente et al. 2019
Spanish amber Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many countries in the Americas **Spanish cuisine **Spanish history **Spanish culture ...
, Early Cretaceous (
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
) Subfamily Coniopteryginae Burmeister 1839 * Tribe Coniopterygini Burmeister 1839 ** †''Paranimboa'' Engel 2016 Burmese amber, Myanmar, Cenomanian * †tribe Phthanoconini Engel 2004 ** †''Phthanoconis'' Engel 2004 Burmese amber, Myanmar, Cenomanian * †''Libanosemidalis'' Azar et al. 2000 Lebanese amber, Barremian * †''Mulleroconis'' Ružičková et al. 2019 Burmese amber, Myanmar, Cenomanian Incertae sedis * †''Apoglaesoconis'' Grimaldi 2000 New Jersey amber, Turonian * †''Cretaconiopteryx'' Liu and Lu 2017 Burmese amber, Myanmar, Cenomanian * †''Jurasiatypus'' Kaddumi 2005
Jordanian amber Jordanian may refer to: * Something of, from, or related to Jordan, a country in the Near East * Jordanian culture * Jordanian people, see Demographics of Jordan * Jordanian cuisine * Jordanian Arabic * Royal Jordanian, an airline See also * L ...
, Albian


References

* (2007): The neuropterid fauna of Dominican and Mexican amber (Neuropterida, Megaloptera, Neuroptera). ''American Museum Novitates'' 3587: 1-58
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:''This article draws heavily on the corresponding article in the German-language Wikipedia.'' {{Authority control Neuroptera Neuroptera families