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Cold Spring Harbor Laboratory (CSHL) is a private, non-profit institution with research programs focusing on cancer, neuroscience, plant biology,
genomics Genomics is an interdisciplinary field of biology focusing on the structure, function, evolution, mapping, and editing of genomes. A genome is an organism's complete set of DNA, including all of its genes as well as its hierarchical, three-dim ...
, and quantitative biology. It is one of 68 institutions supported by the Cancer Centers Program of the U.S. National Cancer Institute (NCI) and has been an NCI-designated Cancer Center since 1987. The Laboratory is one of a handful of institutions that played a central role in the development of molecular genetics and molecular biology. It has been home to eight scientists who have been awarded the Nobel Prize in Physiology or Medicine. CSHL is ranked among the leading basic research institutions in molecular biology and genetics with Thomson Reuters ranking it #1 in the world. CSHL was also ranked #1 in research output worldwide by ''Nature''. The Laboratory is led by Bruce Stillman, a biochemist and cancer researcher. Since its inception in 1890, the institution's campus on the
North Shore North Shore or Northshore may refer to: Geographic features Australia *North Shore (Sydney), a suburban region of Sydney **Electoral district of North Shore **North Shore railway line, Sydney *Noosa North Shore, Queensland * North Shore, New So ...
of
Long Island Long Island is a densely populated island in the southeastern region of the U.S. state of New York (state), New York, part of the New York metropolitan area. With over 8 million people, Long Island is the most populous island in the United Sta ...
has also been a center of biology education. Current CSHL educational programs serve professional scientists, doctoral students in biology, teachers of biology in the K-12 system, and students from the elementary grades through high school. In the past 10 years CSHL conferences & courses have drawn over 81,000 scientists and students to the main campus. For this reason, many scientists consider CSHL a "crossroads of biological science." Since 2009 CSHL has partnered with the
Suzhou Industrial Park The China–Singapore Suzhou Industrial Park (), or Suzhou Industrial Park for short, abbreviated as SIP, is a county-level administrative area located in Suzhou, Jiangsu, China. The industrial park was established in February 1994, as part of th ...
in
Suzhou Suzhou (; ; Suzhounese: ''sou¹ tseu¹'' , Mandarin: ), alternately romanized as Soochow, is a major city in southern Jiangsu province, East China. Suzhou is the largest city in Jiangsu, and a major economic center and focal point of trade ...
, China to create Cold Spring Harbor Asia which annually draws some 3,000 scientists to its meetings and courses. The Cold Spring Harbor Laboratory School of Biological Sciences, formerly the Watson School of Biological Sciences, was founded in 1999. In 2015, CSHL announced a strategic affiliation with the nearby Northwell Health to advance cancer therapeutics research, develop a new clinical cancer research unit at Northwell Health in
Lake Success Lake Success can refer to: *Lake Success (California), lake in California *Lake Success, New York, village in New York *''Lake Success'', a novel by American author Gary Shteyngart Gary Shteyngart (; born July 5, 1972) is a Soviet-born American ...
, NY, to support early-phase clinical studies of new cancer therapies, and recruit and train more clinician-scientists in oncology. CSHL hosts bioRxiv, a preprint repository for publications in the life sciences.


Research programs

Research staff in CSHL's 52 laboratories numbers over 600, including postdoctoral researchers; an additional 125 graduate students and 500 administrative and support personnel bring the total number of employees to over 1,200. Cell biology and genomics
RNA interference (RNAi) and small-RNA biology; DNA replication;
RNA splicing RNA splicing is a process in molecular biology where a newly-made precursor messenger RNA (pre-mRNA) transcript is transformed into a mature messenger RNA (mRNA). It works by removing all the introns (non-coding regions of RNA) and ''splicing'' b ...
; signal transduction; genome structure; non-coding RNAs; deep sequencing;
single-cell sequencing Single-cell sequencing examines the sequence information from individual cells with optimized next-generation sequencing technologies, providing a higher resolution of cellular differences and a better understanding of the function of an individual ...
and analytics;
stem cell In multicellular organisms, stem cells are undifferentiated or partially differentiated cells that can differentiate into various types of cells and proliferate indefinitely to produce more of the same stem cell. They are the earliest type o ...
self-renewal and differentiation; chromatin dynamics; structural biology; advanced
proteomics Proteomics is the large-scale study of proteins. Proteins are vital parts of living organisms, with many functions such as the formation of structural fibers of muscle tissue, enzymatic digestion of food, or synthesis and replication of DNA. In ...
;
mass spectrometry Mass spectrometry (MS) is an analytical technique that is used to measure the mass-to-charge ratio of ions. The results are presented as a ''mass spectrum'', a plot of intensity as a function of the mass-to-charge ratio. Mass spectrometry is use ...
; advanced microscopy. Cancer research
Principal cancer types under study: breast, prostate, blood (leukemia, lymphoma); myelodysplastic syndrome;
melanoma Melanoma, also redundantly known as malignant melanoma, is a type of skin cancer that develops from the pigment-producing cells known as melanocytes. Melanomas typically occur in the skin, but may rarely occur in the mouth, intestines, or eye ( ...
; liver; ovarian and cervical; lung; brain; pancreas. Research foci: drug resistance; cancer
genomics Genomics is an interdisciplinary field of biology focusing on the structure, function, evolution, mapping, and editing of genomes. A genome is an organism's complete set of DNA, including all of its genes as well as its hierarchical, three-dim ...
; tumor microenvironment; cancer metabolism; growth control in mammalian cells; transcriptional and post-transcriptional gene regulation. Neuroscience
Stanley Institute for Cognitive Genomics employs deep sequencing and other tools to study genetics underlying schizophrenia, bipolar disorder, and major depression. Swartz Center for the Neural Mechanisms of Cognition studies cognition in the normal brain as a baseline for understanding dysfunction in psychiatric and neurodegenerative disorders. Other research foci: autism genetics; mapping of the mammalian brain; neural correlates of decision making. Plant biology
Plant genome sequencing; epigenetics and stem cell fate; stem cell signaling; plant-environment interactions; using genetic insights to increase yield of staple crops, e.g., maize, rice, wheat; increase fruit yield in flowering plants, e.g., tomato. Other initiatives: genetics of aquatic plants for biofuel development; lead role in building National Science Foundation's iPlant cyberinfrastructure. Much of this work takes place on 12 acres of farmland at the nearby CSHL Uplands Farm, where expert staff raise crops and ''Arabidopsis'' plants for studies. Seven CSHL faculty members conduct research primarily in plant biology - Drs. David Jackson,
Zachary Lippman Zachary B. Lippman (born 1978) is an American plant biologist and the Jacob Goldfield Professor of Genetics at Cold Spring Harbor Laboratory and a member of the National Academy of Sciences Lippman has used gene editing technology to investigate ...
, Robert Martienssen, Richard McCombie, Ullas Pedmale, Doreen Ware, and
Thomas Gingeras Thomas Raymond Gingeras is an American geneticist and professor at Cold Spring Harbor Laboratory. He is a leader of the National Institutes of Health's ENCODE project. He worked at Affymetrix as Vice President of Biological Sciences before joinin ...
. Simons Center for Quantitative Biology
Genome assembly and validation; mathematical modeling and algorithm development; population genetics; applied statistical and machine learning; biomedical text-mining; computational genomics; cloud computing and Big Data. COVID-19 Scientists at Cold Spring Harbor Laboratory (CSHL), Utah Health University, PEEL Therapeutics, and Weill Cornell Medicine worked to examine the possible function of NETs in COVID-19, gather blood samples from 33 hospitalized individuals, as well as autopsy tissue. Neutrophil Extracellular Traps (NETs) are a form of protection which is utilized by the immune system against certain pathogens.


Educational programs

In addition to its research mission, CSHL has a broad educational mission. The School of Biological Sciences (SBS), established in 1998, awards the Ph.D. degree and fully funds the research program of every student. Students are challenged to obtain their doctoral degree in 4-5 years. The Undergraduate Research Program (URP) for gifted college students (established in 1959), and the Partners for the Future Program for advanced high school students (established in 1990) are now hosted at the SBS. The CSHL Meetings & Courses Program brings over 8,500 scientists from around the world to Cold Spring Harbor annually to share research results – mostly unpublished—in 60 meetings, most held biannually; and to learn new technologies in 30 to 35 professional courses, most offered annually. The Cold Spring Harbor Symposium series, held every year since 1933 with the exception of three years during the Second World War, has been a forum for researchers in genetics, genomics, neuroscience and plant biology. At the Banbury Center, about 25-30 discussion-style meetings are held yearly for a limited number of invited participants. As of 2016, a two-week course at CSHL costs between $3,700 and $4,700 per student and three-day conferences cost about $1,000 per attendee. The DNA Learning Center (DNALC), founded in 1988, was among the early pioneers in developing hands-on genetics lab experiences for middle and high school students. In 2013, 31,000 students on Long Island and New York City were taught genetics labs at the DNALC and satellite facilities in New York. Over 9,000 high school biology teachers have participated in DNALC teacher-training programs. The Cold Spring Harbor Laboratory Press has established a program consisting of seven journals, 190 books, laboratory manuals and protocols, and online services for research preprints.


Funding

In 2015, CSHL had an operating budget of $150 million, over $100 million of which was spent on research. Half of the research budget was devoted to cancer; 25% to neuroscience; 15% to genomics and quantitative biology; and 10% to plant sciences. The sources of research funding in 2015 were: 34% Federal (primarily National Institutes of Health and National Science Foundation); 26% auxiliary activities; 22% private philanthropy; 10% endowment; 3% corporate.


History

The institution took root as The Biological Laboratory in 1890, a summer program for the education of college and high school teachers studying zoology, botany, comparative anatomy and nature. The program began as an initiative of Eugene G. Blackford and Franklin Hooper, director of the Brooklyn Institute of Arts and Sciences, the founding institution of The Brooklyn Museum. In 1904, the Carnegie Institution of Washington established the Station for Experimental Evolution at Cold Spring Harbor on an adjacent parcel. In 1921, the station was reorganized as the Carnegie Institution Department of Genetics. Between 1910 and 1939, the laboratory was the base of the Eugenics Record Office of biologist Charles B. Davenport and his assistant
Harry H. Laughlin Harry Hamilton Laughlin (March 11, 1880 – January 26, 1943) was an American educator and eugenicist. He served as the superintendent of the Eugenics Record Office from its inception in 1910 to its closure in 1939, and was among the most a ...
, two prominent American eugenicists of the period. Davenport was director of the Carnegie Station from its inception until his retirement in 1934. In 1935 the Carnegie Institution sent a team to review the ERO's work, and as a result the ERO was ordered to stop all work. In 1939 the Institution withdrew funding for the ERO entirely, leading to its closure. The ERO's reports, articles, charts, and pedigrees were considered scientific facts in their day, but have since been discredited. Its closure came 15 years after its findings were incorporated into the National Origins Act ( Immigration Act of 1924), which severely reduced the number of immigrants to America from southern and eastern Europe who, Harry Laughlin testified, were racially inferior to the Nordic immigrants from England and Germany. Charles Davenport was also the founder and the first director of the International Federation of Eugenics Organizations in 1925. Today, Cold Spring Harbor Laboratory maintains the full historical records, communications and artifacts of the ERO for historical, teaching and research purposes. The documents are housed in a campus archive and can be accessed online and in a series of multimedia websites. Carnegie Institution scientists at Cold Spring Harbor made many contributions to genetics and medicine. In 1908
George H. Shull George Harrison Shull (April 15, 1874 – September 28, 1954) was an eminent American plant geneticist and the younger brother of botanical illustrator and plant breeder J. Marion Shull. He was born on a farm in Clark County, Ohio, graduated from ...
discovered hybrid corn and the genetic principle behind it called heterosis, or "hybrid vigor." This would become the foundation of modern agricultural genetics. In 1916,
Clarence C. Little Clarence Cook Little (October 6, 1888 – December 22, 1971) was an American genetics, cancer, and tobacco researcher and academic administrator, as well as a eugenicist. Early life C. C. Little was born in Brookline, Massachusetts and attend ...
was among the first scientists to demonstrate a genetic component of cancer. E. Carleton MacDowell in 1928 discovered a strain of mouse called C58 that developed spontaneous leukemia – an early mouse model of cancer. In 1933, Oscar Riddle isolated prolactin, the milk secretion hormone and Wilbur Swingle participated in the discovery of adrenocortical hormone, used to treat Addison's disease.
Milislav Demerec Milislav Demerec (January 11, 1895 – April 12, 1966) was a Croatian-American geneticist, and the director of the Department of Genetics, Carnegie Institution of Washington IW now Cold Spring Harbor Laboratory (CSHL) from 1941 to 1960, re ...
was named director of the Laboratory in 1941. Demerec shifted the Laboratory's research focus to the genetics of microbes, thus setting investigators on a course to study the biochemical function of the gene. During World War Two, Demerec directed efforts at Cold Spring Harbor that resulted in major increases in penicillin production. Beginning in 1941, and annually from 1945, three of the seminal figures of molecular genetics convened summer meetings at Cold Spring Harbor of what they called the Phage Group. Salvador Luria, of Indiana University; Max Delbrück, then of Vanderbilt University; and
Alfred Hershey Alfred Day Hershey (December 4, 1908 – May 22, 1997) was an American Nobel Prize–winning bacteriologist and geneticist. He was born in Owosso, Michigan and received his B.S. in chemistry at Michigan State University in 1930 and his Ph.D. ...
, then of Washington University in St. Louis, sought to discover the nature of genes through study of viruses called bacteriophages that infect bacteria. * In 1945, Delbrück's famous Phage Course was taught for the first time, inspiring, among others, a young James D. Watson; it was repeated for many years after. CSH Symposia important in the cross-fertilization of ideas among molecular biology's pioneers were held in 1951, 1953, 1956, 1961, 1963, and 1966. * At the CSH Symposium in summer 1953, Watson made the first public presentation of DNA's double-helix structure.


Leadership

In 1962, the Department of Genetics, no longer supported by the Carnegie Institution of Washington, formally merged with the Biological Laboratory to form the Cold Spring Harbor Laboratory of Quantitative Biology. In 1970, the name was simplified to Cold Spring Harbor Laboratory. John F. Cairns was appointed as the Director of the merged Cold Spring Harbor Laboratory in 1963 and found that in the absence of continued financial support from the Carnegie Institution of Washington, the newly created institution was in desperate need of funds to support its programs and update facilities. Cairns stabilized the Laboratory and made essential improvements to the facilities. He decided in 1968 that he would step down as Director and he remained at CSHL until 1973, moving then to the Imperial Cancer Research Fund (now Cancer Research UK) in Mill Hill near London, UK. While at CSHL, Cairns performed important experiments on DNA replication in the bacteria ''E. coli''. James D. Watson served as the Laboratory's director and president for 35 years. Upon taking charge in 1968, he focused the Laboratory on cancer research, creating a tumor virus group and successfully obtaining federal funds for an expansion of cancer research capabilities. Watson placed CSHL on a firm financial footing. Inspired by his Nobel collaborator, Francis Crick, Watson initiated a major push to scale-up CSHL research on the brain and psychiatric disorders, beginning in the late 1980s. In 1990, work was completed on the
Arnold and Mabel Beckman Arnold Orville Beckman (April 10, 1900 – May 18, 2004) was an American chemist, inventor, investor, and philanthropist. While a professor at California Institute of Technology, he founded Beckman Instruments based on his 1934 invention of th ...
Laboratory, and the Marks Neuroscience Building was opened in 1999. In 1994, Watson ceased being director of the Laboratory and assumed the title of president. In 2004 he was named chancellor, a position he held until October 2007, when he retired at the age of 79 after views attributed to him on race and intelligence appeared in the British press. In January 2019, CSHL severed all ties with Watson—and revoked his honorary titles—after he unequivocally restated these views in an American Masters television profile. Since 1994 biochemist and cancer biologist Bruce Stillman has led the Laboratory as director, and since 2003 as president. Stillman, a member of the
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nati ...
and a Fellow of the Royal Society, also continues to run a basic research lab, devoted to the study of DNA replication and chromosome maintenance. Stillman is credited with the 1991 discovery and elucidation of the mechanism of the Origin Recognition Complex (ORC), a highly conserved protein complex that recognizes and binds to specific DNA sequences, marking starting points for replication of the entire genome. Stillman has presided over a major expansion of the Laboratory, its size growing threefold since he became director. With construction completed on six linked laboratory buildings on the Hillside Campus in 2009, CSHL added much-needed new laboratory space for cancer and neuroscience research, as well as space for a new program on quantitative biology to bring experts in mathematics, computer science, statistics, and physics to problems in biology.


Notable faculty

*
Douglas Fearon Douglas Thomas Fearon FRS FRCP FMedSci (born 16 October 1942) is an American medical immunologist, who has been since 2003 Sheila Joan Smith Professor of Immunology at the University of Cambridge, a fellow of Trinity College, Cambridge, and ...
, immunologist, Fellow of the Royal Society, member of the 
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nati ...
. *
Leemor Joshua-Tor Leemor Joshua-Tor is the W.M. Keck Professor of Structural Biology at Cold Spring Harbor Laboratory and a Howard Hughes Medical Institute Investigator. Her research focuses on the role of the argonaute complex in RNA interference. Early life and ...
, structural biologist,
Howard Hughes Medical Institute The Howard Hughes Medical Institute (HHMI) is an American non-profit medical research organization based in Chevy Chase, Maryland. It was founded in 1953 by Howard Hughes, an American business magnate, investor, record-setting pilot, engineer, fil ...
 investigator, and
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nati ...
member. *
Adrian R. Krainer Adrian Robert Krainer is a Uruguayan-American biochemist and molecular geneticist known for his research into RNA gene-splicing. Krainer holds the St. Giles Foundation Professorship at Cold Spring Harbor Laboratory in Laurel Hollow, New York. ...
, developed nusinersen to treat
spinal muscular atrophy Spinal muscular atrophy (SMA) is a rare neuromuscular disorder that results in the loss of motor neurons and progressive muscle wasting. It is usually diagnosed in infancy or early childhood and if left untreated it is the most common genetic ...
(SMA), winner of the Breakthrough Prize in Life Sciences and member of the
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nati ...
. * Robert Martienssen, studies epigenetics,
Howard Hughes Medical Institute The Howard Hughes Medical Institute (HHMI) is an American non-profit medical research organization based in Chevy Chase, Maryland. It was founded in 1953 by Howard Hughes, an American business magnate, investor, record-setting pilot, engineer, fil ...
 investigator and Fellow of the Royal Society (FRS). * Bruce Stillman, molecular biologist, Fellow of the Royal Society, Member of the
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nati ...
,
EMBO Embo ( gd, Eurabol, IPA: �iaɾəpɔɫ̪ is a village in the Highland Council Area in Scotland and the former postal county of Sutherland, about north-northeast of Dornoch. On 16 July 1988, Embo declared itself independent from the rest of the ...
and
AAAS AAAS may refer to: * American Academy of Arts and Sciences, a learned society and center for policy research; the publisher of the journal ''Dædalus'' * American Association for the Advancement of Science, an organization that supports scientifi ...
. * Bruce Wallace, geneticist, Member of the
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nati ...
, professor at Virginia Tech * Michael Wigler, genetic engineering of animal cells and molecular biologist, co-discovered the Ras oncogene, Member of the
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nati ...
and
AAAS AAAS may refer to: * American Academy of Arts and Sciences, a learned society and center for policy research; the publisher of the journal ''Dædalus'' * American Association for the Advancement of Science, an organization that supports scientifi ...
. *
Zachary Lippman Zachary B. Lippman (born 1978) is an American plant biologist and the Jacob Goldfield Professor of Genetics at Cold Spring Harbor Laboratory and a member of the National Academy of Sciences Lippman has used gene editing technology to investigate ...
, plant geneticist,
Howard Hughes Medical Institute The Howard Hughes Medical Institute (HHMI) is an American non-profit medical research organization based in Chevy Chase, Maryland. It was founded in 1953 by Howard Hughes, an American business magnate, investor, record-setting pilot, engineer, fil ...
 investigator and winner of a MacArthur Fellowship, commonly referred to as a "Genius grant". *
Christopher Vakoc Christopher Vakoc is a molecular biologist and a professor at Cold Spring Harbor Laboratory. Education Vakoc graduated with a degree in Biochemistry from Pennsylvania State University in 2001. He then attained his M.D. and his Ph.D. from the Uni ...
, leukemia biologist, winner of the Paul Marks Prize for Cancer Research.


Nobel Prize winners

*
Carol Greider Carolyn Widney Greider (born April 15, 1961) is an American molecular biologist and Nobel laureate. She joined the University of California, Santa Cruz as a Distinguished Professor in the department of molecular, cell, and developmental biology ...
, discovered relationship between cellular aging and damage to the ends of chromosomes, called telomeres, in 1992; won Nobel Prize in 2009 with Elizabeth Blackburn and Jack W. Szostak *
Barbara McClintock Barbara McClintock (June 16, 1902 – September 2, 1992) was an American scientist and cytogeneticist who was awarded the 1983 Nobel Prize in Physiology or Medicine. McClintock received her PhD in botany from Cornell University in 1927. There s ...
, discovered transposons ("jumping genes") in 1944; received Nobel Prize in 1983. * Martha Chase and
Alfred Hershey Alfred Day Hershey (December 4, 1908 – May 22, 1997) was an American Nobel Prize–winning bacteriologist and geneticist. He was born in Owosso, Michigan and received his B.S. in chemistry at Michigan State University in 1930 and his Ph.D. ...
, conducted "Waring blender experiments", confirmed DNA as the genetic material in 1952; Hershey was awarded the Nobel Prize with Salvador Luria and Max Delbrück in 1969 *
Richard J. Roberts Sir Richard John Roberts (born 6 September 1943) is a British biochemist and molecular biology, molecular biologist. He was awarded the 1993 Nobel Prize in Physiology or Medicine with Phillip Allen Sharp for the discovery of introns in eukaryo ...
and Phillip A. Sharp shared a Nobel in 1993 for the discovery of discontinuous, or "split" genes, which revealed the
RNA splicing RNA splicing is a process in molecular biology where a newly-made precursor messenger RNA (pre-mRNA) transcript is transformed into a mature messenger RNA (mRNA). It works by removing all the introns (non-coding regions of RNA) and ''splicing'' b ...
mechanism. * James D. Watson, shared a Nobel Prize with
Francis Crick Francis Harry Compton Crick (8 June 1916 – 28 July 2004) was an English molecular biologist, biophysicist, and neuroscientist. He, James Watson, Rosalind Franklin, and Maurice Wilkins played crucial roles in deciphering the helical struc ...
and
Maurice Wilkins Maurice Hugh Frederick Wilkins (15 December 1916 – 5 October 2004) was a New Zealand-born British biophysicist and Nobel laureate whose research spanned multiple areas of physics and biophysics, contributing to the scientific understanding o ...
in 1962 for their discovery of the double helix structure of DNA. From 1968 to 1993 Watson served as the CSHL Director. In 2007 the CSHL suspended him for his support for scientific racism but after he issued a public apology he was allowed to retain honorary titles, though he was relieved his leadership and managerial roles. In 2019 CSHL rescinded his honorary titles after he made public remarks again suggesting IQ and race were related, comments which CSHL viewed as a reversal of the apology he gave in 2007. In 2020 the Watson School of Biological Sciences (WSBS) was renamed to delete any reference to him.


Double Helix Medal

Each year, Cold Spring Harbor Laboratory awards a Double Helix Medal to individuals who have positively impacted human health, either by raising awareness and funds for biomedical research, or contributing in a major way to biological and biomedical research.


See also

*
Salk Institute for Biological Studies The Salk Institute for Biological Studies is a scientific research institute located in the La Jolla community of San Diego, California, U.S. The independent, non-profit institute was founded in 1960 by Jonas Salk, the developer of the polio vacci ...
* Whitehead Institute * Massachusetts Institute of Technology * Harvard University *
Rockefeller University The Rockefeller University is a private biomedical research and graduate-only university in New York City, New York. It focuses primarily on the biological and medical sciences and provides doctoral and postdoctoral education. It is classif ...
* California Institute of Technology *
The Scripps Research Institute Scripps Research, previously known as The Scripps Research Institute (TSRI), is a nonprofit American medical research facility that focuses on research and education in the biomedical sciences. Headquartered in San Diego, California, the institu ...


References


External links

*
Dolan DNA Learning CenterEugenics ArchiveCold Spring Harbor Laboratory Press
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