Cola (plant)
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''Cola'' is a genus of
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s native to the
tropical forest Tropical forests are forested ecoregions with tropical climates – that is, land areas approximately bounded by the Tropic of Cancer, tropics of Cancer and Tropic of Capricorn, Capricorn, but possibly affected by other factors such as prevailing ...
s of
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
, classified in the family
Malvaceae Malvaceae (), or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include Theobroma cacao, cacao, Cola (plant), cola, cotton, okra, Hibiscus sabdariffa, ...
, subfamily
Sterculioideae Sterculioideae is a subfamily of the family (biology), family Malvaceae containing evergreen and deciduous tree and shrub genera, that were previously placed in the obsolete family Sterculiaceae. Taxonomy A 2006 molecular study indicated the Ste ...
(previously in the separate family
Sterculiaceae Sterculiaceae was a family of flowering plant based on the genus '' Sterculia''. Genera formerly included in Sterculiaceae are now placed in the family Malvaceae, in the subfamilies: Byttnerioideae, Dombeyoideae, Helicteroideae and Sterculioide ...
). Species in this genus are sometimes referred to as kola tree or
kola nut The kola nut ( Yoruba: ''obi'', Dagbani: ''guli'', Hausa: goro, Igbo: ''ọjị'', Sängö: ''gôro,'' Swahili: ''mukezu'') is the seed of certain species of plant of the genus ''Cola'', placed formerly in the cocoa family Sterculiaceae and ...
for the
caffeine Caffeine is a central nervous system (CNS) stimulant of the methylxanthine chemical classification, class and is the most commonly consumed Psychoactive drug, psychoactive substance globally. It is mainly used for its eugeroic (wakefulness pr ...
-containing fruit produced by the trees that is often used as a flavoring ingredient in beverages. The genus was thought to be closely related to the
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
n genus '' Theobroma'', or cocoa, but the latter is now placed in a different subfamily. They are
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
trees, growing up to 20 m tall (about 60 feet), with glossy ovoid
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
up to 30 cm long and star-shaped fruit.


Origin and distribution

''Cola'' is a genus of the Family
Malvaceae Malvaceae (), or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include Theobroma cacao, cacao, Cola (plant), cola, cotton, okra, Hibiscus sabdariffa, ...
with approximately 100 to 125 species occurring in the evergreen lowland and montane forest of continental (primarily tropical) Africa. The earliest known evidence of ''Cola'' is ''Cola amharaensis'', a well-preserved fossil leaf compression from the late Oligocene Guang River flora of Ethiopia and dated to 27.23 ± 0.1 Ma.
Kola nut The kola nut ( Yoruba: ''obi'', Dagbani: ''guli'', Hausa: goro, Igbo: ''ọjị'', Sängö: ''gôro,'' Swahili: ''mukezu'') is the seed of certain species of plant of the genus ''Cola'', placed formerly in the cocoa family Sterculiaceae and ...
s are seeds harvested from pods, primarily from the species ''
Cola nitida ''Cola nitida'' is a species of plant belonging to the family Malvaceae. It is a tree native to the rainforests of tropical West Africa. Common names include kola nut, cola, kola and bitter kola. The seeds contain caffeine and are chewed as a s ...
'' and ''
Cola acuminata ''Cola acuminata'' is a species in the genus ''Cola'', of the family Malvaceae, native to tropical Africa. It is generally known for its fruit, the kola nut, originally used to impart the cola flavor in manufactured beverages, such as Coca-Col ...
''. Outside mainland
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
, some species are cultivated for their nuts in
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
,
Jamaica Jamaica is an island country in the Caribbean Sea and the West Indies. At , it is the third-largest island—after Cuba and Hispaniola—of the Greater Antilles and the Caribbean. Jamaica lies about south of Cuba, west of Hispaniola (the is ...
and elsewhere in the humid tropics.


Species

* ''
Cola acuminata ''Cola acuminata'' is a species in the genus ''Cola'', of the family Malvaceae, native to tropical Africa. It is generally known for its fruit, the kola nut, originally used to impart the cola flavor in manufactured beverages, such as Coca-Col ...
'' (P.Beauv.) Schott & Endl. * '' Cola alba'' A.Chev. * '' Cola altissima'' Engl. * '' Cola angustifolia'' K.Schum. * '' Cola anomala'' K.Schum. * '' Cola argentea'' Mast. * '' Cola attiensis'' Aubrév. & Pellegr. * '' Cola ballayi'' Cornu ex Heckel * '' Cola bilenguensis'' Pellegr. * '' Cola bipindensis'' Engl. * '' Cola bodardii'' Pellegr. * ''
Cola boxiana Cola is a carbonated soft drink flavored with vanilla, cinnamon, citrus oils, and other flavorings. Cola became popular worldwide after the American pharmacist John Stith Pemberton invented Coca-Cola, a trademarked brand, in 1886, which was imi ...
'' Brenan & Keay * '' Cola brevipes'' K.Schum. * '' Cola bruneelii'' De Wild. * '' Cola buesgenii'' Engl. * '' Cola buntingii'' Baker f. * '' Cola cabindensis'' Exell * '' Cola caricifolia'' (G.Don) K.Schum. * '' Cola cauliflora'' Mast. * '' Cola cecidiifolia'' Cheek * '' Cola chlamydantha'' K.Schum. * '' Cola chlorantha'' F.White * '' Cola clavata'' Mast. * '' Cola coccinea'' Engl. & K.Krause * '' Cola congolana'' De Wild. & T.Durand * '' Cola cordifolia'' (Cav.) R.Br. * '' Cola crispiflora'' K.Schum. * '' Cola digitata'' Mast. * '' Cola discoglypremnophylla'' Brenan & A.P.D.Jones * '' Cola diversifolia'' De Wild. & T.Durand * '' Cola duparquetiana'' Baill. * '' Cola edeensis'' Engl. & K.Krause * '' Cola elegans'' Pierre ex Breteler * '' Cola fibrillosa'' Engl. & K.Krause * '' Cola ficifolia'' Mast. * '' Cola flaviflora'' Engl. & K.Krause * '' Cola flavovelutina'' K.Schum. * '' Cola gabonensis'' Mast. * '' Cola gigantea'' A.Chev. * '' Cola gigas'' Baker f. * '' Cola gilgiana'' Engl. * '' Cola gilletii'' De Wild. * '' Cola glabra'' Brenan & Keay * '' Cola glaucoviridis'' Pellegr. * '' Cola greenwayi'' Brenan * '' Cola griseiflora'' De Wild. * '' Cola heterophylla'' (P.Beauv.) Schott & Endl. * '' Cola hispida'' Brenan & Keay * '' Cola hypochrysea'' K.Schum. * '' Cola idoumensis'' Pellegr. * '' Cola kimbozensis'' Cheek * '' Cola lasiantha'' Engl. & K.Krause * '' Cola lateritia'' K.Schum. * '' Cola laurifolia'' Mast. * '' Cola le-testui'' Pellegr. * '' Cola lepidota'' K.Schum. * '' Cola letouzeyana'' Nkongmeneck * '' Cola liberica'' Jongkind * '' Cola lissachensis'' Pellegr. * '' Cola lizae'' N.Hallé * '' Cola lomensis'' Engl. & K.Krause * '' Cola lorougnonis'' Aké Assi * '' Cola louisii'' Germ. * '' Cola lukei'' Cheek * '' Cola macrantha'' K.Schum. * '' Cola mahoundensis'' Pellegr. * '' Cola marsupium'' K.Schum. * '' Cola mayimbensis'' Pellegr. * '' Cola mayumbensis'' Exell * '' Cola megalophylla'' Brenan & Keay * '' Cola metallica'' Cheek * '' Cola millenii'' K.Schum. * '' Cola minor'' Brenan * '' Cola mixta'' A.Chev. * '' Cola mossambicensis'' Wild * '' Cola mosserayana'' Germ. * '' Cola nana'' Engl. & K.Krause * '' Cola natalensis'' Oliv. * '' Cola ndongensis'' Engl. & K.Krause * '' Cola nigerica'' Brenan & Keay * ''
Cola nitida ''Cola nitida'' is a species of plant belonging to the family Malvaceae. It is a tree native to the rainforests of tropical West Africa. Common names include kola nut, cola, kola and bitter kola. The seeds contain caffeine and are chewed as a s ...
'' (Vent.) Schott & Endl. * '' Cola noldeae'' Exell & Mendonça * '' Cola obtusa'' Engl. & K.Krause * '' Cola octoloboides'' Brenan * '' Cola pachycarpa'' K.Schum. * '' Cola pallida'' A.Chev. * '' Cola philipi-jonesii'' Brenan & Keay * '' Cola pierlotii'' Germ. * '' Cola porphyrantha'' Brenan * '' Cola praeacuta'' Brenan & Keay * '' Cola pseudoclavata'' Cheek * '' Cola pulcherrima'' Engl. * '' Cola quentinii'' Cheek * '' Cola quintasii'' Engl. * '' Cola reticulata'' A.Chev. * '' Cola ricinifolia'' Engl. & K.Krause * '' Cola rondoensis'' Cheek * '' Cola rostrata'' K.Schum. * '' Cola ruawaensis'' Cheek * '' Cola rubra'' A.Chev. * '' Cola scheffleri'' K.Schum. * '' Cola sciaphila'' Louis ex Germ. * '' Cola selengana'' Germ. * '' Cola semecarpophylla'' K.Schum. * '' Cola simiarum'' Sprague ex Brenan & Keay * '' Cola sphaerocarpa'' A.Chev. * '' Cola sphaerosperma'' Heckel * '' Cola stelechantha'' Brenan * '' Cola subglaucescens'' Engl. * '' Cola suboppositifolia'' Cheek * '' Cola sulcata'' Engl. * '' Cola tessmannii'' Engl. & K.Krause * '' Cola triloba'' (R.Br.) K.Schum. * '' Cola tsandensis'' Pellegr. * '' Cola uloloma'' Brenan * '' Cola umbratilis'' Brenan & Keay * '' Cola urceolata'' K.Schum. * '' Cola usambarensis'' Engl. * '' Cola vandersmisseniana'' Germ. * '' Cola verticillata'' (Thonn.) Stapf ex A.Chev. * '' Cola welwitschii'' Exell & Mendonça ex R.Germ. * '' Cola winkleri'' Engl.


Gallery

Produits forestiers non ligneux récoltés à Tayap.jpg, ''Cola pachycarpa'' Cola gigantea at Kepong Botanical Garden 20230625 104415.jpg, ''Cola gigantea'' Cola natalensis 5Dsr 4440.jpg, ''Cola natalensis''


References

{{Authority control Malvaceae genera