A chronospecies is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is physically, morphologically, and/or genetically distinct from the original ancestors. Throughout the change, there is only one species in the lineage at any point in time, as opposed to cases where divergent evolution produces contemporary species with a common ancestor. The related term paleospecies (or palaeospecies) indicates an extinct species only identified with

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

material. That identification relies on distinct similarities between the earlier fossil specimens and some proposed descendant although the exact relationship to the later species is not always defined. In particular, the range of variation within all the early fossil specimens does not exceed the observed range that exists in the later species. A paleosubspecies (or palaeosubspecies) identifies an extinct

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

that evolved into the currently-existing form. The connection with relatively-recent variations, usually from the

Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...

, often relies on the additional information available in subfossil material. Most of the current species have changed in size and so adapted to the climatic changes during the last ice age (see Bergmann's Rule). The further identification of fossil specimens as part of a "chronospecies" relies on additional similarities that more strongly indicate a specific relationship with a known species. For example, relatively recent specimens, hundreds of thousands to a few million years old with consistent variations (such as always smaller but with the same proportions) as a living species might represent the final step in a chronospecies. The possible identification of the immediate ancestor of the living

taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...

may also rely on stratigraphic information to establish the age of the specimens. The concept of chronospecies is related to the phyletic gradualism model of evolution, and it also relies on an extensive fossil record since morphological changes accumulate over time, and two very different organisms could be connected by a series of intermediaries.

Examples

* ''

Bison A bison (: bison) is a large bovine in the genus ''Bison'' (from Greek, meaning 'wild ox') within the tribe Bovini. Two extant taxon, extant and numerous extinction, extinct species are recognised. Of the two surviving species, the American ...

'' (several paleospecies and -subspecies) * Marine sloths (paleospecies) * '' Coragyps'' (chronospecies) * '' Gymnogyps'' (paleospecies) * ''

Panthera ''Panthera'' is a genus within the family (biology), family Felidae, and one of two extant genera in the subfamily Pantherinae. It contains the largest living members of the cat family. There are five living species: the jaguar, leopard, lion, ...

'' (numerous chrono- and paleospecies and -subspecies) * '' Valdiviathyris'' (no visible change since the

Priabonian The Priabonian is, in the ICS's geologic timescale, the latest age or the upper stage of the Eocene Epoch or Series. It spans the time between . The Priabonian is preceded by the Bartonian and is followed by the Rupelian, the lowest stage ...

, 35 million years ago)

References

External links

{{Wiktionary Evolutionary biology Biostratigraphy Phylogenetics

Examples

See also

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