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Anagenesis
Anagenesis is the gradual evolution of a species that continues to exist as an interbreeding population. This contrasts with cladogenesis, which occurs when branching or splitting occurs, leading to two or more lineages and resulting in separate species. Anagenesis does not always lead to the formation of a new species from an ancestral species. When speciation does occur as different lineages branch off and cease to interbreed, a core group may continue to be defined as the original species. The evolution of this group, without extinction or species selection, is anagenesis. Hypotheses One hypothesis is that during the speciation event in anagenetic evolution, the original populations increases quickly, and then racks up genetic variation over long periods of time by mutation and recombination in a stable environment. Other factors, such as selection or genetic drift will have such a significant effect on genetic material and physical traits that a species can be acknowl ...
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Cladogenesis
Cladogenesis is an evolutionary splitting of a parent species into two distinct species, forming a clade. This event usually occurs when a few organisms end up in new, often distant areas or when environmental changes cause several extinctions, opening up ecological niches for the survivors and causing population bottlenecks and founder effects changing allele frequencies of diverging populations compared to their ancestral population. The events that cause these species to originally separate from each other over distant areas may still allow both of the species to have equal chances of surviving, reproducing, and even evolving to better suit their environments while still being two distinct species due to subsequent natural selection, mutations and genetic drift. Cladogenesis is in contrast to anagenesis, in which an ancestral species gradually accumulates change, and eventually, when enough is accumulated, the species is sufficiently distinct and different enough from its ...
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Australopithecus Anamensis
''Australopithecus anamensis'' is a hominin species that lived roughly between 4.3 and 3.8 million years ago, and is the oldest known ''Australopithecus'' species, Nearly 100 fossil specimens of ''A. anamensis'' are known from Kenya and Ethiopia, representing over 20 individuals. The first fossils of ''A. anamensis'' discovered are dated to around 3.8 and 4.2 million years ago and were found in Kanapoi and Allia Bay in northern Kenya. '' A. afarensis'' is normally accepted to have emerged within this lineage. However, ''A. anamensis'' and ''A. afarensis'' appear to have lived side-by-side for at least some period of time, and whether the lineage that led to extant humans emerged in ''A. afarensis'', or directly in ''A. anamensis'' is not fully settled. Fossil evidence determines that ''Australopithecus anamensis'' is the earliest hominin species in the Turkana Basin, but likely co-existed with ''afarensis'' towards the end of its existence. ''A. anamensis'' and ''A. afarensis'' ...
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Punctuated Equilibrium
In evolutionary biology, punctuated equilibrium (also called punctuated equilibria) is a Scientific theory, theory that proposes that once a species appears in the fossil record, the population will become stable, showing little evolution, evolutionary change for most of its geological history. : ''Reprinted in'' * * This state of little or no morphological change is called ''stasis''. When significant evolutionary change occurs, the theory proposes that it is generally restricted to rare and geologic time scale, geologically rapid events of branching speciation called cladogenesis. Cladogenesis is the process by which a species splits into two distinct species, rather than one species gradually transforming into another. Punctuated equilibrium is commonly contrasted with phyletic gradualism, the idea that evolution generally occurs uniformly by the steady and gradual transformation of whole lineages (anagenesis). In 1972, paleontologists Niles Eldredge and Stephen Jay Goul ...
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Paraphyly
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancest ...
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Chronospecies
A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is physically, morphologically, and/or genetically distinct from the original ancestors. Throughout the change, there is only one species in the lineage at any point in time, as opposed to cases where divergent evolution produces contemporary species with a common ancestor. The related term paleospecies (or palaeospecies) indicates an extinct species only identified with fossil material. That identification relies on distinct similarities between the earlier fossil specimens and some proposed descendant although the exact relationship to the later species is not always defined. In particular, the range of variation within all the early fossil specimens does not exceed the observed range that exists in the later species. A paleosubspecies (o ...
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Species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. Other ways of defining species include their karyotype, DNA sequence, morphology (biology), morphology, behaviour, or ecological niche. In addition, palaeontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses) are given a binomial nomenclature, two-part name, a "binomen". The first part of a binomen is the name of a genus to which the species belongs. The second part is called the specific name (zoology), specific name or the specific ...
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Evolution
Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, resulting in certain characteristics becoming more or less common within a population over successive generations. The process of evolution has given rise to biodiversity at every level of biological organisation. The scientific theory of evolution by natural selection was conceived independently by two British naturalists, Charles Darwin and Alfred Russel Wallace, in the mid-19th century as an explanation for why organisms are adapted to their physical and biological environments. The theory was first set out in detail in Darwin's book ''On the Origin of Species''. Evolution by natural selection is established by observable facts about living organisms: (1) more offspring are often produced than can possibly survive; (2) phenotypic variatio ...
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Extinction
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and recover. As a species' potential Range (biology), range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxon, Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the Fossil, fossil record) after a period of apparent absence. Over five billion species are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryotes globally, possibly many times more if microorganisms are included. Notable extinct animal species include Dinosaur, non-avian dinosaurs, Machairodontinae, saber-toothed cats, and mammoths. Through evolution, species arise through the process of specia ...
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Speciation
Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within lineages. Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book ''On the Origin of Species''. He also identified sexual selection as a likely mechanism, but found it problematic. There are four geographic modes of speciation in nature, based on the extent to which speciating populations are isolated from one another: allopatric speciation, allopatric, peripatric speciation, peripatric, parapatric speciation, parapatric, and sympatric speciation, sympatric. Whether genetic drift is a minor or major contributor to speciation is the subject of much ongoing discussion. Rapid sympatric speciation can take place through polyploidy, such as by doubling of chromosome number; the result is progeny wh ...
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Marc Ereshefsky
Marc Ereshefsky is a professor of philosophy at the University of Calgary, specializing in the philosophy of science and the philosophy of biology. His research focuses on issues on the intersection of philosophy and biology. Ereshefsky is specifically known for his work on taxonomy, systematics and natural kinds. His research has been supported by the Social Sciences and Humanities Research Council of Canada, the Canadian Institutes of Health Research, and the National Science Foundation The U.S. National Science Foundation (NSF) is an Independent agencies of the United States government#Examples of independent agencies, independent agency of the Federal government of the United States, United States federal government that su .... Personal life Ereshefsky lives in Calgary with his wife, two sons, Jacob and Josh, and their puppy. His favourite weekend activities include hiking in the nearby Rocky Mountains, and going for runs. Publications Books * ''The Units of Evolution ...
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Taxonomy (biology)
In biology, taxonomy () is the science, scientific study of naming, defining (Circumscription (taxonomy), circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxon, taxa (singular: taxon), and these groups are given a taxonomic rank; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain (biology), domain, kingdom (biology), kingdom, phylum (''division'' is sometimes used in botany in place of ''phylum''), class (biology), class, order (biology), order, family (biology), family, genus, and species. The Swedish botanist Carl Linnaeus is regarded as the founder of the current system of taxonomy, having developed a ranked system known as Linnaean taxonomy for categorizing organisms. With advances in the theory, data and analytical technology of biological systematics, the Linnaean system has transfo ...
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Plio-Pleistocene
The Plio-Pleistocene is an informally described geological pseudo-period, which begins about 5 million years ago (Mya) and, drawing forward, combines the time ranges of the formally defined Pliocene and Pleistocene epochs—marking from about 5 Mya to about 12 kya. Nominally, the Holocene epoch—the last 12 thousand years—would be excluded, but most Earth scientists would probably treat the current times as incorporated into the term "Plio-Pleistocene"; see below. In the contexts of archaeology, paleontology, and paleoanthropology, the Plio-Pleistocene is a very useful period to which scientists may assign the long and continuous run in East Africa of datable sedimentary layers and their contents (e.g. the Bouri Formation). These contents collectively present a focused view of the continuous evolution of the region's large vertebrates, especially the evolution of some African apes ( hominids) to the earliest hominins; and then the development of the early humans and ...
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