Chanaresuchus
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''Chanaresuchus'' is an extinct
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of proterochampsid archosauriform. It was of modest size for a proterochampsian, being on average just over a meter in length. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Chanaresuchus bonapartei'' was named in 1971. Its fossils were found in from the early
Carnian The Carnian (less commonly, Karnian) is the lowermost stage (stratigraphy), stage of the Upper Triassic series (stratigraphy), Series (or earliest age (geology), age of the Late Triassic Epoch (reference date), Epoch). It lasted from 237 to 227.3 ...
-age Chañares Formation in La Rioja Province,
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
. ''Chanaresuchus'' appears to be one of the most common archosauriforms from the Chañares Formation due to the abundance of specimens referred to the genus. Much of the material has been found by the La Plata-Harvard expedition of 1964–65. ''Chanaresuchus'' is the most well-described proterochampsid in the subfamily Rhadinosuchinae. A second proposed species, ''C. ischigualastensis'', was named in 2012 from the late Carnian-age Ischigualasto Formation, was briefly assigned to ''Chanaresuchus'' before being moved to its own genus '' Pseudochampsa'' in 2014. ''C. bonapartei'' has been reported from the Santa Maria Formation in Brazil, but the Brazilian fossils were given their own genus '' Kuruxuchampsa'' in 2023.


Description

''Chanaresuchus'' has a low, elongate skull that is characteristic of proterochampsians. The skull is quite broad posteriorly with a narrow snout, varying in length from around 165mm to 260mm in the largest individuals. The nares are slit-like and positioned away from the tip of the
rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit ** podium * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ...
, farther up the skull. The
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammals h ...
is slightly down-curved. The skull table is highly ornamented in larger specimens, with the
dermal bone A dermal bone or investing bone or membrane bone is a bony structure derived from intramembranous ossification forming components of the vertebrate skeleton, including much of the skull, jaws, gill covers, shoulder girdle, fin rays ( lepidotrich ...
s well sculptured. The palate of ''Chanaresuchus'' has two elongate
choana The choanae (: choana), posterior nasal apertures or internal nostrils are two openings found at the back of the nasal passage between the nasal cavity and the pharynx, in humans and other mammals (as well as crocodilians and most skinks). They ...
e. Two small openings anterior to the choanae may be anterior palatine foramina that could have been used for access to
vomeronasal organ The vomeronasal organ (VNO), or Jacobson's organ, is the paired auxiliary olfactory (smell) sense organ located in the soft tissue of the nasal septum, in the nasal cavity just above the roof of the mouth (the hard palate) in various tetrapods ...
s. The
secondary palate The secondary palate is an anatomical structure that divides the nasal cavity from the oral cavity in many vertebrates. In human embryology, it refers to that portion of the hard palate that is formed by the growth of the two palatine shelves med ...
formed between these two sets of openings may have been an adaptation for breathing through the snout while underwater. Unlike other proterochampsians and early archosaurs, ''Chanaresuchus'' had little body armor. The only
osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amph ...
s found are small and scale-like, forming a single row down the back. They run from the neck to the hip, ending at the last presacral vertebra. They most likely continue down the tail, although tail osteoderms are not preserved in any specimens. There are roughly three osteoderms overlying each vertebra. The foot of ''Chanaresuchus'' differs from other related archosaurs in that the inner toes are inlarged, whereas other primitive archosaurs retain a more symmetrical pattern. The first digit is reduced but robust, the second digit is the thickest, and the third digit is the longest, although somewhat slender in comparison to the others. The fourth digit is very slim and the fifth consists of only a
metatarsal The metatarsal bones or metatarsus (: metatarsi) are a group of five long bones in the midfoot, located between the tarsal bones (which form the heel and the ankle) and the phalanges ( toes). Lacking individual names, the metatarsal bones are ...
spur.


Paleoecology

A semiaquatic lifestyle similar to phytosaurs and modern day
crocodilia Crocodilia () is an order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorph pseudosuchia ...
ns has been proposed for ''Chanaresuchus'', as is suggested by the secondary palate and upward facing orbits and nostrils. However, some evidence, such as a lack of aquatic
amphibian Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s found from the Chañares Formation, suggests that the area was relatively dry during the time of deposition. A terrestrial lifestyle is possible, since the osteoderm structure of ''Chanaresuchus'' is compact, more similar to terrestrial squamates than to crocodilians. The
depositional environment In geology, depositional environment or sedimentary environment describes the combination of physical, chemical, and biological processes associated with the deposition of a particular type of sediment and, therefore, the rock types that will b ...
of the locality from which specimens of ''Chanaresuchus'' have been found was in close proximity to an area of high
volcanic A volcano is commonly defined as a vent or fissure in the crust of a planetary-mass object, such as Earth, that allows hot lava, volcanic ash, and gases to escape from a magma chamber below the surface. On Earth, volcanoes are most often fo ...
activity, because it was in an active rift basin. It is possible that all of the recovered specimens had died in a single event of mass mortality and may have been buried on a
fluvial A river is a natural stream of fresh water that flows on land or inside caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of its course if it ru ...
strandline. The mortality-causing event was most likely linked to regional volcanic activity. The locality from which specimens of ''C. bonapartei'' have been found is well known for its abundance of tetrapods. Theraspids include the
dicynodont Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivores that typically bore a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, t ...
'' Dinodontosaurus'', and
cynodonts Cynodontia () is a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extinct ances ...
such as ''
Probainognathus ''Probainognathus'' meaning “progressive jaw” is an extinct genus of cynodonts that lived around 235 to 221.5 million years ago, during the Late Triassic in what is now Argentina. Together with the genus ''Bonacynodon'' from Brazil, ''Probain ...
'' and '' Massetognathus'', the latter being the most abundant taxon of the locality. Archosaurs were less common, though highly diverse.
Ornithodira Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. Di ...
ns include ''
Lewisuchus ''Lewisuchus'' is a genus of archosaur that lived during the Late Triassic (early Carnian). As a silesaurid dinosauriform, it was a member of the group of reptiles most commonly considered to be the closest relatives of dinosaurs (possibly tru ...
'' (= '' Pseudolagosuchus''), ''
Lagerpeton ''Lagerpeton'' is a genus of lagerpetid avemetatarsalian, comprising a single species, ''L. chanarensis''. First described from the Chañares Formation of Argentina by A. S. Romer in 1971, ''Lagerpeton'' anatomy is somewhat incompletely k ...
'', and ''
Marasuchus ''Marasuchus'' (meaning "Mara (mammal), Mara crocodile") is a genus of basal Dinosauriformes, dinosauriform archosaur which is possibly synonymous with ''Lagosuchus''. Both genera lived during the Late Triassic in what is now La Rioja Province, A ...
''. Other archosaurs include ''
Gracilisuchus ''Gracilisuchus'' (meaning "slender crocodile") is an extinct genus of tiny pseudosuchian (a group which includes the ancestors of crocodilians) from the Late Triassic of Argentina. It contains a single species, ''G. stipanicicorum'', which is pl ...
'' and ''
Luperosuchus ''Luperosuchus'' (meaning "vexing" or "difficult crocodile") is an extinct genus of loricatan pseudosuchian reptile (historically known as a "rauisuchian") which contains only a single species, ''Luperosuchus fractus.'' It is known from the Cha ...
''. Another proterochampsian, which was named alongside ''Chanaresuchus'' in 1971, is '' Gualosuchus''. It is very similar in appearance to ''Chanaresuchus'', differing only in size and cranial proportions.


References


External links


''Chanaresuchus''
in the
Paleobiology Database The Paleobiology Database (PBDB) is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database originated in the NCEAS-funded Phanerozoic Marine Pale ...
{{Taxonbar, from=Q744348 Proterochampsia Prehistoric reptile genera Middle Triassic reptiles of South America Late Triassic reptiles of South America Triassic Argentina Fossils of Argentina Santa Maria Formation Triassic Brazil Fossils of Brazil Fossil taxa described in 1971 Taxa named by Alfred Romer