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Flower chafers are a group of scarab beetles comprising the subfamily Cetoniinae. Many species are diurnal and visit
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s for
pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
and
nectar Nectar is a viscous, sugar-rich liquid produced by Plant, plants in glands called nectaries, either within the flowers with which it attracts pollination, pollinating animals, or by extrafloral nectaries, which provide a nutrient source to an ...
, or to browse on the petals. Some species also feed on
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
. The group is also called fruit and flower chafers, flower beetles and flower scarabs. There are around 4,000 species, many of them still undescribed. Ten tribes are presently recognized: Cetoniini, Cremastocheilini, Diplognathini, Goliathini, Gymnetini, Phaedimini, Schizorhinini, Stenotarsiini, Taenioderini, and Xiphoscelidini. The former tribes Trichiini and Valgini were elevated in rank to subfamily. The tribe Gymnetini is the biggest of the American tribes, and Goliathini contains the largest species, and is mainly found in the
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
regions of
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
.


Description

Adult flower chafers are usually brightly coloured beetles, often metallic, and somewhat flattened in shape. The insertions of the antennae are visible from above, while the mandibles and labrum are hidden by the clypeus. The elytra lack a narrow membranous margin and are truncate to expose the pygidium. The abdominal spiracles are diverging so that several lie on the abdominal sternites with at least one exposed. The fore coxae are conical and produced ventrally, while the mid coxae are transverse or only slightly oblique. The mesothoracic epimera is visible from above. The tarsi are each equipped with a pair of simple (not forked) tarsal claws of subequal size. A feature possessed by adults of many flower chafers, especially Cetoniini, is lateral emargination of the elytra. Larvae are stout-bodied and very hairy with short legs. The head is partly covered by the prothorax. Each antenna has the apical segment as wide as the penultimate segment. The galea and lacinia are used to form a mala. The anal cleft is transverse. The mandible has a ventral stridulating area. The labrum is symmetrical with a deeply pigmented notch on each side of the midline.


Biology

Adult cetoniines are herbivorous, being found on flowers (from which they consume nectar and pollen), tree sap and rotting fruit. Larvae generally live and feed in decaying plant matter (including decaying wood) or soil. In captivity, cetoniine larvae will feed on soft fruit. Many species in the tribe Cremastocheilini are known to be predaceous, feeding on hymenopteran larvae or soft-bodied
nymphs A nymph (; ; sometimes spelled nymphe) is a minor female nature deity in ancient Greek folklore. Distinct from other Greek goddesses, nymphs are generally regarded as personifications of nature; they are typically tied to a specific place, land ...
of Auchenorrhyncha. '' Spilophorus'' spp. have been noted feeding on the nesting material and excrement of South African passerine birds, while '' Spilophorus maculatus'' has been recorded feeding on '' Oxyrhachis'' sp. nymphs and '' Hoplostomus fuligineus'' is known to feed on the brood of honey bees in South Africa and the pupae of the wasp '' Belonogaster petiolata''. '' Campsiura javanica'' feeds on the larvae of '' Ropalidia montana'' in southern India. '' Cremastocheilus stathamae'' feeds on ants of the genus '' Myrmecocystus''. In terms of movement, adults are considered some of the best flyers among beetles. They can hover above and land on flowers or fruit. When threatened by predators, they escape by either performing a rush take off or by falling toward the ground and then flying before impact. Many cetoniines fly with their elytra closed, as their hindwings can unfold and slide out under the elytra during flight (thanks to the emargination of the elytra). Larvae of some taxa can crawl on their backs using their tergal folds, which are covered in strong bristles. Others crawl on their legs.


Systematics and taxonomy

The following list contains the genera and subtribes in ten tribes of subfamily Cetoniinae, according to Catalogue of Life and Scarabaeidae of the World (2023).


Cetoniini

Authority: Leach, 1815 * Subtribe Cetoniina Leach, 1815 # '' Aethiessa'' Burmeister, 1842 # '' Anatona'' Burmeister, 1842 # '' Anelaphinis'' Kolbe, 1892 # '' Aphelinis'' Antoine, 1987 # '' Atrichelaphinis'' Kraatz, 1898 # '' Atrichiana'' Distant, 1911 # '' Badizoblax'' Thomson, 1877 # '' Centrantyx'' Fairmaire, 1884 # '' Cetonia'' Fabricius, 1775 # '' Chewia'' Legrand, 2004 # '' Chiloloba'' Burmeister, 1842 # '' Cosmesthes'' Kraatz, 1880 # '' Cosmiophaena'' Kraatz, 1898 # '' Dischista'' Burmeister, 1842 # '' Dolichostethus'' Kolbe, 1892 # '' Elaphinis'' Burmeister, 1842 # '' Enoplotarsus'' Lucas, 1859 # '' Erlangeria'' Preiss, 1902 # '' Gametis'' Burmeister, 1842 # '' Gametoides'' Antoine, 2005 # '' Glycosia'' Schoch, 1896 # '' Glycyphana'' Burmeister, 1842 # '' Gymnophana'' Arrow, 1910 # '' Hemiprotaetia'' Mikšić, 1963 # '' Heterocnemis'' Albers, 1852 # '' Heterotephraea'' Antoine, 2002 # '' Lorkovitschia'' Mikšić, 1968 # '' Marmylida'' Thomson, 1880 # '' Mireia'' Ruter, 1953 # '' Niphobleta'' Kraatz, 1880 # '' Pachnoda'' Burmeister, 1842 # '' Pachnodoides'' Alexis & Delpont, 2002 # '' Pachytephraea'' De Palma & Malec, 2020 # '' Paleopragma'' Thomson, 1880 # '' Paranelaphinis'' Antoine, 1988 # '' Paraprotaetia'' Moser, 1907 # '' Pararhabdotis'' Kraatz, 1899 # '' Parastraella'' Antoine, 2005 # '' Paraxeloma'' Holm, 1988 # '' Parelaphinis'' Marais & Holm, 1989 # '' Phaneresthes'' Kraatz, 1894 # '' Phonotaenia'' Kraatz, 1880 # '' Phoxomeloides'' Schoch, 1898 # '' Podopholis'' Moser, 1915 # '' Podopogonus'' Moser, 1917 # '' Pogonopus'' Arrow, 1910 # '' Polystalactica'' Kraatz, 1882 # '' Protaetia'' Burmeister, 1842 # '' Protaetiomorpha'' Mikšić, 1968 # '' Pseudotephraea'' Kraatz, 1882 # '' Reineria'' Mikšić, 1968 # '' Rhabdotis'' Burmeister, 1842 # '' Rhabdotops'' Krikken, 1981 # '' Rhyxiphloea'' Burmeister, 1842 # '' Ruteraetia'' Krikken, 1980 # '' Simorrhina'' Kraatz, 1886 # '' Somalibia'' Lansberge, 1882 # '' Stalagmosoma'' Burmeister, 1842 # '' Systellorrhina'' Kraatz, 1895 # '' Tephraea'' Burmeister, 1842 # '' Thyreogonia'' Reitter, 1898 # '' Trichocephala'' Moser, 1916 # '' Tropinota'' Mulsant, 1842 # '' Walsternoplus'' Allsopp, Jákl & Rey, 2023 # '' Xeloma'' Kraatz, 1881 * Subtribe Euphoriina Horn, 1880 # '' Chlorixanthe'' Bates, 1889 # '' Euphoria'' Burmeister, 1842 * Subtribe Leucocelina Kraatz, 1882 # '' Acrothyrea'' Kraatz, 1882 # '' Alleucosma'' Schenkling, 1921 # '' Amaurina'' Kolbe, 1895 # '' Analleucosma'' Antoine, 1989 # '' Cyrtothyrea'' Kolbe, 1895 # '' Discopeltis'' Burmeister, 1842 # '' Grammopyga'' Kolbe, 1895 # '' Heteralleucosma'' Antoine, 1989 # '' Homothyrea'' Kolbe, 1895 # '' Leucocelis'' Burmeister, 1842 # '' Lonchothyrea'' Kolbe, 1895 # '' Mausoleopsis'' Lansberge, 1882 # '' Mecaspidiellus'' Antoine, 1997 # '' Molynoptera'' Kraatz, 1897 # '' Molynopteroides'' Antoine, 1989 # '' Oxythyrea'' Mulsant, 1842 # '' Paleira'' Reiche, 1871 # '' Paralleucosma'' Antoine, 1989 # '' Phoxomela'' Schaum, 1844 # '' Pseudalleucosma'' Antoine, 1989 # '' Pseudooxythyrea'' Baraud, 1985


Cremastocheilini

Authority: Burmeister & Schaum, 1841 * Subtribe Aspilina Krikken, 1984 # '' Aspilochilus'' Rojkoff, 2013 # '' Aspilus'' Westwood, 1874 # '' Protochilus'' Krikken, 1976 * Subtribe Coenochilina Burmeister, 1842 # '' Arielina'' Rossi, 1958 # '' Astoxenus'' Péringuey, 1907 # '' Basilewskynia'' Schein, 1957 # '' Coenochilus'' Schaum, 1841 * Subtribe Cremastocheilina Burmeister & Schaum, 1841 # '' Centrochilus'' Krikken, 1976 # '' Clinterocera'' Motschulsky, 1857 # '' Cremastocheilus'' Knoch, 1801 # '' Cyclidiellus'' Krikken, 1976 # '' Cyclidinus'' Westwood, 1874 # '' Cyclidius'' MacLeay, 1838 # '' Genuchinus'' Westwood, 1874 # '' Lissomelas'' Bates, 1889 # '' Paracyclidius'' Howden, 1971 # '' Platysodes'' Westwood, 1874 # '' Psilocnemis'' Burmeister, 1842 * Subtribe Cymophorina Krikken, 1984 # '' Cymophorus'' Kirby, 1827 # '' Myrmecochilus'' Wasmann, 1900 # '' Rhagopteryx'' Burmeister, 1842 * Subtribe Genuchina Krikken, 1984 # '' Genuchus'' Kirby, 1825 # '' Meurguesia'' Ruter, 1969 # '' Problerhinus'' Deyrolle, 1864 * Subtribe Goliathopsidina Krikken, 1984 # '' Goliathopsis'' Janson, 1881 * Subtribe Heterogeniina Krikken, 1984 # '' Heterogenius'' Moser, 1911 # '' Pseudastoxenus'' Bourgoin, 1921 * Subtribe Lissogeniina Krikken, 1984 # '' Chthonobius'' Burmeister, 1847 # '' Lissogenius'' Schaum, 1844 * Subtribe Macromina Burmeister & Schaum, 1842 # '' Brachymacroma'' Kraatz, 1896 # '' Campsiura'' Hope, 1831 # '' Macromina'' Westwood, 1874 # '' Pseudopilinurgus'' Moser, 1918 * Subtribe Nyassinina Krikken, 1984 # '' Nyassinus'' Westwood, 1879 * Subtribe Oplostomina Krikken, 1984 # '' Anatonochilus'' Péringuey, 1907 # '' Oplostomus'' MacLeay, 1838 # '' Placodidus'' Péringuey, 1900 # '' Scaptobius'' Schaum, 1841 * Subtribe Pilinurgina Krikken, 1984 # '' Callynomes'' Mohnike, 1873 # '' Centrognathus'' Guérin-Méneville, 1840 # '' Parapilinurgus'' Arrow, 1910 # '' Periphanesthes'' Kraatz, 1880 # '' Pilinurgus'' Burmeister, 1842 # '' Priska'' Jákl, 2018 * Subtribe Spilophorina Krikken, 1984 # '' Spilophorus'' Westwood, 1848 * Subtribe Telochilina Krikken, 1984 # '' Telochilus'' Krikken, 1975 * Subtribe Trichoplina Krikken, 1984 # '' Lecanoderus'' Kolbe, 1907 # '' Trichoplus'' Burmeister, 1842 * Subtribe Trogodina Krikken, 1984 # '' Pseudoscaptobius'' Krikken, 1976 # '' Trogodes'' Boheman, 1857


Diplognathini

Authority: Burmeister, 1842 # '' Anoplocheilus'' MacLeay, 1838 # '' Anthracophora'' Burmeister, 1842 # '' Anthracophorides'' Moser, 1918 # '' Apocnosoides'' Antoine, 2001 # '' Charadronota'' Burmeister, 1842 # '' Conradtia'' Kolbe, 1892 # '' Diphrontis'' Gerstaecker, 1882 # '' Diplognatha'' Gory & Percheron, 1833 # '' Eriulis'' Burmeister, 1842 # '' Hadrodiplognatha'' Kraatz, 1898 # '' Heteropseudinca'' Valck Lucassen, 1933 # '' Lamellothyrea'' Krikken, 1980 # '' Metallopseudinca'' Valck Lucassen, 1933 # '' Niphetophora'' Kraatz, 1883 # '' Nselenius'' (Holm & Perissinotto, 2004) # '' Odontorrhina'' Burmeister, 1842 # '' Parapoecilophila'' Hauser, 1904 # '' Phonopleurus'' Moser, 1919 # '' Pilinopyga'' Kraatz, 1888 # '' Porphyronota'' Burmeister, 1842 # '' Pseudinca'' Kraatz, 1880 # '' Stethopseudinca'' Valck Lucassen, 1933 # '' Tetragonorrhina'' Kraatz, 1896 # ''
Trichostetha ''Trichostetha'' (Greek 'hairy' + 'chest') is a genus of beetle in the Scarabaeidae, scarab beetle family. It is endemic to southern Africa, and its species most commonly occur in mountainous terrain. The genus includes several species that have ...
'' Burmeister, 1842 # '' Triplognatha'' Krikken, 1987 # '' Trymodera'' Gerstaecker, 1867 # '' Uloptera'' Burmeister, 1842


Goliathini

Authority: Latreille, 1829 * Subtribe Dicronocephalina Krikken, 1984 # '' Dicronocephalus'' # '' Platynocephalus'' * Subtribe Goliathina Latreille, 1829 # '' Fornasinius'' Bertoloni, 1852 # '' Goliathus'' Lamarck, 1801 # '' Hegemus'' Thomson, 1881 # '' Hypselogenia'' Burmeister, 1840 * Subtribe Ichnestomina Burmeister, 1842 # '' Gariep'' Péringuey, 1907 # '' Ichnestoma'' Gory & Percheron, 1833 # '' Karooida'' Perissinotto, 2020 # '' Mzansica'' Perissinotto, 2020 * Subtribe Rhomborhinina Westwood, 1842 # '' Anagnathocera'' Arrow, 1922 # '' Anisorrhina'' Westwood, 1842 # '' Asthenorhella'' Westwood, 1874 # '' Asthenorhina'' Westwood, 1843 # '' Bietia'' Fairmaire, 1898 # '' Caelorrhina'' Hope, 1841 # '' Cheirolasia'' Westwood, 1842 # '' Chloresthia'' Fairmaire, 1905 # '' Chlorocala'' Kirby, 1828 # '' Chondrorrhina'' Kraatz, 1880 # '' Compsocephalus'' White, 1845 # '' Cosmiomorpha'' Saunders, 1852 # '' Cyphonocephalus'' Westwood, 1842 # '' Desfontainesia'' Alexis & Delpont, 1999 # '' Dicellachilus'' Waterhouse, 1905 # '' Dicheros'' Gory & Percheron, 1833 # '' Dicronorhina'' Hope, 1837 # '' Diphyllomorpha'' Hope, 1843 # '' Dymusia'' Burmeister, 1842 # '' Euchloropus'' Arrow, 1907 # '' Eudicella'' White, 1839 # '' Eutelesmus'' Waterhouse, 1880 # '' Gnathocera'' Kirby, 1825 # '' Gnorimimelus'' Kraatz, 1880 # '' Hemiheterorrhina'' Mikšić, 1974 # '' Herculaisia'' Seilliere, 1910 # '' Heterorhina'' Westwood, 1842 # '' Ingrisma'' Fairmaire, 1893 # '' Ischnoscelis'' Burmeister, 1842 # '' Jumnos'' Saunders, 1839 # '' Lansbergia'' Ritsema, 1888 # '' Lophorrhina'' Westwood, 1842 # '' Lophorrhinides'' Perissinotto, Clennell & Beinhundner, 2019 # '' Mawenzhena'' Alexis & Delpont, 2001 # '' Mecynorhina'' Hope, 1837 # '' Moseriana'' Ruter, 1965 # '' Mystroceros'' Burmeister, 1842 # ''
Narycius ''Narycius opalus'' is a flower chafer beetle that is endemic to the Western Ghats of India. It is the sole species in the genus. The adult male has a prominent projection on the head, while the female has a much shorter horn. This structure is p ...
'' Dupont, 1835 # '' Neomystroceros'' Alexis & Delpont, 1998 # '' Neophaedimus'' Lucas, 1870 # '' Neoscelis'' Schoch, 1897 # '' Pedinorrhina'' Kraatz, 1880 # '' Plaesiorrhina'' Westwood, 1842 # '' Petrovitzia'' Mikšić, 1965 # '' Priscorrhina'' Krikken, 1984 # '' Pseudodiceros'' Mikšić, 1974 # '' Pseudotorynorrhina'' Mikšić, 1967 # '' Ptychodesthes'' Kraatz, 1883 # '' Raceloma'' Thomson, 1877 # '' Rhamphorrhina'' Klug, 1855 # '' Rhinarion'' Ruter, 1965 # '' Rhomborhina'' Hope, 1837 # '' Scythropesthes'' Kraatz, 1880 # '' Smicorhina'' Westwood, 1847 # '' Spelaiorrhina'' Lansberge, 1886 # '' Stephanorrhina'' Burmeister, 1842 # '' Taurhina'' Burmeister, 1842 # '' Tmesorrhina'' Westwood, 1841 # '' Torynorrhina'' Arrow, 1907 # '' Trichoneptunides'' Legrand, 2001 # '' Trigonophorinus'' Pouillaude, 1913 # '' Trigonophorus'' Hope, 1831


Gymnetini

Authority: Kirby, 1827 * Subtribe Blaesiina Schoch, 1895 # '' Blaesia'' Burmeister, 1842 # '' Halffterinetis'' Morón & Nogueira, 2007 * Subtribe Gymnetina Kirby, 1827 # '' Allorrhina'' Burmeister, 1842 # '' Amazula'' Kraatz, 1882 # '' Amithao'' Thomson, 1878 # '' Argyripa'' Thomson, 1878 # '' Astroscara'' Schürhoff, 1937 # '' Badelina'' Thomson, 1880 # '' Balsameda'' Thomson, 1880 # '' Chiriquibia'' Bates, 1889 # '' Clinteria'' Burmeister, 1842 # '' Clinteroides'' Schoch, 1898 # '' Cotinis'' Burmeister, 1842 - (Green June Beetles) # '' Desicasta'' Thomson, 1878 # '' Guatemalica'' Neervoort Van De Poll, 1886 # '' Gymnephoria'' Ratcliffe, 2019 # '' Gymnetina'' Casey, 1915 # '' Gymnetis'' MacLeay, 1819 # '' Hadrosticta'' Kraatz, 1892 # '' Heterocotinis'' Martinez, 1948 # '' Hologymnetis'' Martinez, 1949 # '' Hoplopyga'' Thomson, 1880 # '' Hoplopygothrix'' Schürhoff, 1933 # '' Howdenypa'' Arnaud, 1993 # '' Jansonia'' Schürhoff, 1937 # '' Macrocranius'' Schürhoff, 1935 # '' Madiana'' Ratcliffe & Romé, 2019 # '' Marmarina'' Kirby, 1827 # '' Neocorvicoana'' Ratcliffe & Mico, 2001 # '' Pseudoclinteria'' Kraatz, 1882 # '' Stethodesma'' Bainbridge, 1840 # '' Tiarocera'' Burmeister, 1842


Phaedimini

Authority: Schoch, 1894 # '' Hemiphaedimus'' Mikšić, 1972 # '' Phaedimus'' Waterhouse, 1841 # '' Philistina'' MacLeay, 1838 # '' Theodosia'' Thomson, 1880


Schizorhinini

Authority: Burmeister, 1842 * Subtribe Lomapterina Burmeister, 1842 # '' Agestrata'' Eschscholtz, 1829 # '' Ischiopsopha'' Gestro, 1874 # '' Lomaptera'' Gory & Percheron, 1833 # '' Macronota'' Hoffmannsegg, 1817 # '' Megaphonia'' Schürhoff, 1933 # '' Morokia'' Janson, 1905 # '' Mycterophallus'' Neervoort Van De Poll, 1886 # '' Thaumastopeus'' Kraatz, 1885 * Subtribe Schizorhinina Burmeister, 1842 # '' Anacamptorrhina'' Blanchard, 1842 # '' Aphanesthes'' Kraatz, 1880 # '' Aurum'' Hutchinson & Moeseneder, 2019 # '' Axillonia'' Krikken, 2018 # '' Bisallardiana'' Antoine, 2003 # '' Chalcopharis'' Heller, 1902 # '' Chlorobapta'' Kraatz, 1880 # '' Chondropyga'' Kraatz, 1880 # '' Clithria'' Burmeister, 1842 # '' Diaphonia'' Newman, 1840 # '' Dichrosoma'' Kraatz, 1885 # '' Digenethle'' Thomson, 1877 # '' Dilochrosis'' Thomson, 1878 # '' Eupoecila'' Burmeister, 1842 # '' Grandaustralis'' Hutchinson & Moeseneder, 2013 # '' Hemichnoodes'' Kraatz, 1880 # '' Hemipharis'' Burmeister, 1842 # '' Kerowagia'' Delpont, 1996 # '' Lenosoma'' MacLeay, 1863 # '' Lyraphora'' Kraatz, 1880 # '' Macrotina'' Strand, 1934 # '' Metallesthes'' Kraatz, 1880 # '' Microdilochrosis'' Jákl, 2009 # '' Microlomaptera'' Kraatz, 1885 # '' Micropoecila'' Kraatz, 1880 # '' Navigator'' Moeseneder & Hutchinson, 2016 # '' Neoclithria'' Neervoort Van De Poll, 1886 # '' Neorrhina'' Thomson, 1878 # '' Octocollis'' Moeseneder & Hutchinson, 2012 # '' Panglaphyra'' Kraatz, 1880 # '' Peotoxus'' Krikken, 1983 # '' Poecilopharis'' Kraatz, 1880 # '' Pseudoclithria'' Neervoort Van De Poll, 1886 # '' Rigoutorum'' Hutchinson & Moeseneder, 2022 # '' Schizorhina'' Kirby, 1825 # '' Schochidia'' Berg, 1898 # '' Stenopisthes'' Moser, 1913 # '' Storeyus'' Hasenpusch & Moeseneder, 2010 # '' Tafaia'' Valck Lucassens, 1939 # '' Tapinoschema'' Thomson, 1880 # '' Territonia'' Krikken, 2018 # '' Trichaulax'' Kraatz, 1880


Stenotarsiini

Authority: Kraatz, 1880 * Subtribe Anochiliina Krikken, 1984 # '' Anochilia'' Burmeister, 1842 # '' Epistalagma'' Fairmaire, 1880 * Subtribe Chromoptiliina Krikken, 1984 # '' Chromoptilia'' Westwood, 1842 # '' Descarpentriesia'' Ruter, 1964 * Subtribe Coptomiina Schenkling, 1921 # '' Bricoptis'' Burmeister, 1842 # '' Coptomia'' Burmeister, 1842 # '' Coptomiopsis'' Pouillaude, 1919 # '' Eccoptomia'' Kraatz, 1880 # '' Euchilia'' Burmeister, 1842 # '' Euryomia'' Burmeister, 1842 # '' Heterocranus'' Bourgoin, 1919 # '' Hiberasta'' Fairmaire, 1901 # '' Hyphelithia'' Kraatz, 1880 # '' Liostraca'' Burmeister, 1842 # '' Micreuchilia'' Pouillaude, 1917 # '' Micropeltus'' Blanchard, 1842 # '' Pareuchilia'' Kraatz, 1880 # '' Pygora'' Burmeister, 1842 # '' Pyrrhopoda'' Kraatz, 1880 # '' Vieuella'' Ruter, 1964 * Subtribe Doryscelina Schenkling, 1921 # '' Doryscelis'' Burmeister, 1842 # '' Epixanthis'' Burmeister, 1842 # '' Hemiaspidius'' Krikken, 1982 # '' Pararhynchocephala'' Paulian, 1991 # '' Parepixanthis'' Kraatz, 1893 # '' Pseudepixanthis'' Kraatz, 1880 # '' Rhynchocephala'' Fairmaire, 1883 * Subtribe Euchroeina Paulian & Descarpentries, 1982 # '' Euchroea'' Burmeister, 1842 * Subtribe Heterophanina Schoch, 1894 # '' Heterophana'' Burmeister, 1842 # '' Oxypelta'' Pouillaude, 1920 # '' Pogoniotarsus'' Kraatz, 1880 # '' Pogonotarsus'' Burmeister, 1842 # '' Zebinus'' Fairmaire, 1894 * Subtribe Heterosomatina Krikken, 1984 # '' Heterosoma'' Schaum, 1844 # '' Plochilia'' Fairmaire, 1896 * Subtribe Pantoliina Krikken, 1984 # '' Bonoraella'' Ruter, 1978 # '' Celidota'' Burmeister, 1842 # '' Cyriodera'' Burmeister, 1842 # '' Dirrhina'' Burmeister, 1842 # '' Hemilia'' Kraatz, 1880 # '' Lucassenia'' Olsoufieff, 1940 # '' Melanchroea'' Kraatz, 1900 # '' Moriaphila'' Kraatz, 1880 # '' Pantolia'' Burmeister, 1842 # '' Tetraodorhina'' Blanchard, 1842 * Subtribe Parachiliina Krikken, 1984 # '' Parachilia'' Burmeister, 1842 * Subtribe Stenotarsiina Kraatz, 1880 # '' Callipechis'' Burmeister, 1842 # '' Ischnotarsia'' Kraatz, 1880 # '' Rhadinotaenia'' Kraatz, 1900 # '' Stenotarsia'' Burmeister, 1842 # '' Vadonidella'' Ruter, 1973


Taenioderini

Authority: Mikšić, 1976 * Subtribe Chalcotheina Mikšić, 1976 # '' Anocoela'' Moser, 1914 # '' Chalcothea'' Burmeister, 1842 # '' Chalcotheomima'' Mikšić, 1970 # '' Clerota'' Burmeister, 1842 # '' Glyptothea'' Bates, 1889 # '' Glyptotheomima'' Mikšić, 1976 # '' Hemichalcothea'' Mikšić, 1970 # '' Microchalcothea'' Moser, 1910 # '' Paraplectrone'' Mikšić, 1985 # '' Penthima'' Kraatz, 1892 # '' Plectrone'' Wallace, 1867 # '' Pseudochalcothea'' Ritsema, 1882 # '' Pseudochalcotheomima'' Mikšić, 1985 * Subtribe Taenioderina Mikšić, 1976 # '' Bacchusia'' Mikšić, 1976 # '' Bombodes'' Westwood, 1848 # '' Carneluttia'' Mikšić, 1976 # '' Coilodera'' Hope, 1831 # '' Costinota'' Schürhoff, 1933 # '' Eumacronota'' Mikšić, 1976 # '' Euremina'' Westwood, 1867 # '' Euselates'' Thomson, 1880 # '' Gnorimidia'' Lansberge, 1887 # '' Ixorida'' Thomson, 1880 # '' Macronotops'' Krikken, 1977 # '' Meroloba'' Thomson, 1880 # '' Pleuronota'' Kraatz, 1892 # '' Stenonota'' Fairmaire, 1889 # '' Taeniodera'' Burmeister, 1842 # '' Xenoloba'' Bates, 1889


Xiphoscelidini

Authority: Krikken, 1984 # '' Aporecolpa'' Lansberge, 1886 # '' Callophylla'' Moser, 1916 # '' Heteroclita'' Burmeister, 1842 # '' Ischnostomiella'' Krikken, 1978 # '' Meridioclita'' Krikken, 1982 # '' Myodermidius'' Bourgoin, 1920 # '' Neoclita'' Perissinotto, 2017 # '' Oroclita'' Krikken, 1982 # '' Plochiliana'' Ruter, 1978 # '' Protoclita'' Krikken, 1978 # '' Rhinocoeta'' Burmeister, 1842 # '' Scheinia'' Ruter, 1957 # '' Xiphoscelis'' Burmeister, 1842 # '' Xiphosceloides'' Holm, 1992


References

*Evans, Arthur W
Generic Guide to New World Scarabs
*Orozco, Jesus
American Cetoniinae
*Sakai, K. & S. Nagai, 1998. ''The cetoniine Beetles of the Wor''ld. Tokyo: Mushi-Sha. 421p Excellent illustrations of most species. *White, Richard E. (1998). ''Beetles : A Field Guide to the Beetles of North America''. Boston: Houghton Mifflin. .


External links




Illustrated key to South Asian chafers

Punctate Flower Chafer Factfile


Gallery

African Cetoniinae website. Literature page.
flickr
tagged images.

by Oz Rittner {{DEFAULTSORT:Flower Chafer Scarabaeidae Polyphaga subfamilies