The Cerro Bandera Formation is a
geological formation
A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics (lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock expo ...
in
Neuquén Province,
Argentina
Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
, in northern
Patagonia
Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and ...
, which dates to the Early Miocene, around 21 to 17.5 million years ago. It predominantly consists of
pyroclastic
Pyroclastic rocks (derived from the el, πῦρ, links=no, meaning fire; and , meaning broken) are clastic rocks composed of rock fragments produced and ejected by explosive volcanic eruptions. The individual rock fragments are known as pyroc ...
deposits, which were deposited in a
semi-arid
A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi- ...
environment. It is divided up into a number of members. The diverse fauna of the Cerro Bandera Formation include a variety of turtles and birds, also includes many mammals such as
South American native ungulates
South American native ungulates, commonly abbreviated as SANUs, are extinct ungulate-like mammals of controversial affinities that were indigenous to South America prior to the Great American Biotic Interchange. They comprise five major groups ...
(
notoungulates,
litopterns,
astrapotheres
Astrapotheria is an extinct order of South American and Antarctic hoofed mammals that existed from the late Paleocene to the Middle Miocene, ."The uruguaytheriine Astrapotheriidae from the rich middle Miocene Honda Group of the upper Magdalena R ...
) as well as
armadillo
Armadillos (meaning "little armored ones" in Spanish) are New World placental mammals in the order Cingulata. The Chlamyphoridae and Dasypodidae are the only surviving families in the order, which is part of the superorder Xenarthra, al ...
s, and
caviomorph rodents.
[ ]
Material was copied from this source, which is available under a Creative Commons Attribution 3.0 International License
The Cerro Bandera Formation comprises a series of isolated outcrops that represent the relicts of an old alluvial filling developed on small local valleys. It is composed of a succession of reworked pyroclastic deposits alternating with primary pyroclastic and scant sandstone levels. These deposits were originally recognized at the northeast of Barda Negra, south of Cerro Bandera and northwest of Sierra del Portezuelo; new exposures are herein recognised to the northwestern slope of Cerro Bayo Mesa, Neuquén Province, Argentina. The recovered fossil remains correspond to birds of the family
Falconidae
The falcons and caracaras are around 60 species of diurnal birds of prey that make up the family Falconidae (representing all extant species in the order Falconiformes). The family is divided into three subfamilies, Herpetotherinae, which incl ...
and 17 families of mammals, among which are remarkable ''
Cramauchenia'', ''
Proadinotherium
''Proadinotherium'' is an extinct genus of toxodontid. It lived between the Late Oligocene and the Early Miocene in what is now South America.
Description
This genus is only known from very partial remains. From the comparison with its relative ...
'', ''
Eosteiromys'', and ''
Caviocricetus'', plus a species of ''
Protypotherium'' with more primitive dentition than those known from the
Santacrucian
The Santacrucian age is a period of geologic time (17.5 – 16.3 Ma) within the Early Miocene epoch of the Neogene, used more specifically with SALMA classification in South America. It follows the Colhuehuapian and precedes the Friasian age.
...
SALMA. This association confirms a Colhuehuapian SALMA (Early Miocene) for this unit. This fauna is markedly different from that recorded from the lower section of the
Chichinales Formation
The Chichinales Formation is a geological formation in Río Negro Province, Argentina which dates from the Late Oligocene to the Early Miocene, around 23 to 17.5 million years ago. It predominantly consists of pyroclastic deposits, which were depo ...
, Río Negro Province. The degree of faunistic differentiation between these probably synchronous units could be a result of local palaeoenvironmental differences.
The fossiliferous localities of the Cerro Bandera Formation are latitudinally equivalent to that of the
Chichinales Formation
The Chichinales Formation is a geological formation in Río Negro Province, Argentina which dates from the Late Oligocene to the Early Miocene, around 23 to 17.5 million years ago. It predominantly consists of pyroclastic deposits, which were depo ...
. However, there are marked differences between their known faunal compositions. The only genera shared by both units are ''
Cladosictis'', ''
Cramauchenia'', ''
Proeutatus'', ''
Protypotherium'', and ''
Stenotatus
''Stenotatus'' is an extinct genus of cingulate, belonging to the family Dasypodidae. It lived from the Early to the Late Miocene in South America.
Description
Stenotatus was a small to medium-sized armadillo, not exceeding 4 kilograms in weigh ...
''. To this must be added the recent discovery of remains of ''
Eoviscaccia
''Eoviscaccia'' is an extinct genus of chinchillid rodent that lived during the Early Oligocene (Tinguirirican) to the Early Miocene (Colhuehuapian) in what is now South America. Fossils of this genus have been found in the Cerro Bandera, Chich ...
'' in the Chichinales Formation. In this unit the remains of turtles,
hegetotheriid and
leontiniid notoungulates are extremely abundant; while that in the Cerro Bandera Formation there are still no known remains of turtles, the interatheriids predominate over the hegetotheriids and the large herbivorous ungulates are mostly represented by the astrapotheres. In addition, the presence of at least two new genera only known from the Cerro Bandera Formation belonging to the families
Astrapotheriidae and
Cephalomyidae
Cephalomyidae is an extinct family of caviomorph rodents from South America. The specific relationships of the family are uncertain, and affinities to both chinchilloid and cavioid rodents have been supported. Most recently, Kramarz in 2005 per ...
, infers a certain degree of faunal differentiation with respect not only to the Chichinales Formation, but also to the typical Colhuehuapian localities of the central Patagonia.
Stratigraphy
The Cerro Bandera Formation occurs as a succession of small isolated whitish to yellowish-brown outcrops, aligned along ancient depressions and paleo-valleys carved in Late Cretaceous and Paleocene sediments. In the study area, the deposits of this unit overlie in erosive discordance several units of the Neuquén Group, including the
Huincul,
Cerro Lisandro and
Portezuelo Formations, from the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
, as well as over
Danian
The Danian is the oldest age or lowest stage of the Paleocene Epoch or Series, of the Paleogene Period or System, and of the Cenozoic Era or Erathem. The beginning of the Danian (and the end of the preceding Maastrichtian) is at the Creta ...
marine sediments of the
Roca Formation in the
Malargüe Group.
The known fossiliferous deposits in the Cerro Bandera Formation consist of isolated concentrations of fossil remains partially exposed on the surface due to the effect of rainwater and wind action. In general, fossil remains tend to be found in greater abundance in those sectors where the sequence is less altered. Possibly, the same factors that contributed to the alteration and devitrification of the
pyroclastites also influenced the degradation of the bone materials. In this sense, the dental pieces seem to be the most resistant, having been found in some cases parts of dental series in correct position within the sediment, although any bone remains were completely missing.
At the type locality of the Cerro Bandera Formation, the succession consists of dominantly chonitic (consolidated pyroclastic deposits of very fine grain size, less than 0.062 mm,) and
tuffitic deposits. In this outcrop were found two fossiliferous horizons within were found within deposits corresponding to strongly pedoturbed tuffaceous chonites (
paleosol
In the geosciences, paleosol (''palaeosol'' in Great Britain and Australia) is an ancient soil that formed in the past. The precise definition of the term in geology and paleontology is slightly different from its use in soil science.
In geol ...
s). The lower fossiliferous horizon corresponds to the basal deposits of this unit, where remains of small vertebrates were found together with freshwater gastropods.
On the northeastern slope of Barda Negra, the Cerro Bandera Formation exhibits a granocreccive sequence, with predominantly
bentonitic (lacustrine) deposits at the base, gradually grading towards the top to deposits with higher epiclastic content (tuffites with sandy lenses), where fluvial sandstones with conglomeratic lenses bearing silicified logs dominate. In this locality, the basal bentonite deposits are commercially exploited in several mining properties, from where some dental and bone remains have been extracted, together with inadequately preserved fossil logs.
In the locality of Sierra del Portezuelo Norte, the unit is formed by an alternation of tuffites, bentonite chonites and tuffaceous chonites. At least four fossiliferous horizons have been recognized at the same locality, of which only the third horizon, located approximately 21 m from the base of the profile, has contributed fossiliferous material significant enough for this analysis. This horizon is located in pedoturbed deposits of tuffaceous chonites, which contributed numerous mammal remains along with scarce, poorly preserved beetle nests.
Finally, a fourth fossiliferous locality has been found on the northwestern slope of Cerro Bayo Mesa, a site where the Cerro Bandera Formation had not been previously recognized. This unit occurs there as a dominantly chonitic and tuffitic succession of thin thickness, with a single fossiliferous horizon, located ca. 13 m from the base of the profile, in levels of
pedogenized tuffaceous chonites.''
Paleofauna
Reptiles
Testudines
Birds
Mammals
Meridiungulates
Xenarthrans
Rodents
Metatherians
References
{{Reflist
Geologic formations of Argentina
Colhuehuapian
Burdigalian
Aquitanian (stage)
Miocene Series of South America
Neogene Argentina
Tuff formations
Sandstone formations
Fluvial deposits
Lacustrine deposits
Deltaic deposits
Fossiliferous stratigraphic units of South America
Paleontology in Argentina
Formations
Geology of Neuquén Province
Geology of Patagonia