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''Caucasotachea vindobonensis'' is a
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of medium-sized air-breathing land snail, a
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate gastropod in the family
Helicidae Helicidae is a large, diverse family of small to large, air-breathing land snails, sometimes called the "typical snails." A number of species in this family are valued as food items, including ''Cornu aspersum'' (formerly ''Helix aspersa'') the ...
. The scientific name is derived from the Celtic settlement
Vindobona Vindobona (from Gaulish ''windo-'' "white" and ''bona'' "base/bottom") was a Roman military camp on the site of the modern city of Vienna in Austria. The settlement area took on a new name in the 13th century, being changed to Berghof, or now si ...
, now known as
Vienna en, Viennese , iso_code = AT-9 , registration_plate = W , postal_code_type = Postal code , postal_code = , timezone = CET , utc_offset = +1 , timezone_DST ...
, the capital of Austria. This species was assigned to the genus '' Cepaea'' for a long time. However, molecular biologist research showed, that this species is not closely related to ''Cepaea'', but belongs instead to the genus ''Caucasotachea''.


Description

The right-coiled, globular shell of ''C. vindobonensis'' is about 17–21 mm high and 20–25 mm broad with 5,5-6 whorls. The umbilicus is completely covered in adult specimens. The lip is always red-brown at the inner and outer side with a fine white stripe at its margin. This specific lip coloration allows to separate it from the conchological similar '' Cepaea hortensis'' (pure white lip) and '' Cepaea nemoralis'' (consistent dark brown lip), with which ''C. vindobonensis'' often co-occurs. The shell is whitish or yellowish, with about 5 brown stripes, the upper two are usually weak. The lowermost stripe is quite near to the covered umbilicus. There is a slight variation of shell colour within this species (see below). The animal itself is yellowish with grey tentacles. Examples of variation in shell colour of ''C. vindobonensis'': File:Cepaea_vindobonensis_01.JPG File:Cepaea_vindobonensis_02.JPG File:Cepaea_vindobonensis_03.JPG


Distribution and habitat

The distribution of this species is
Pontic Pontic, from the Greek ''pontos'' (, ), or "sea", may refer to: The Black Sea Places * The Pontic colonies, on its northern shores * Pontus (region), a region on its southern shores * The Pontic–Caspian steppe, steppelands stretching from n ...
, Pannonian and Balkanian. It occurs in Albania,
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
,
Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Mac ...
, Bosnia and Herzegovina, Croatia,
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. Th ...
(near threatened (NT) Its westernmost, but non-indigenous, record is from Western Bohemia.), Germany ( reintroduced), Greece),
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croa ...
, Italy, Latvia, Macedonia, Moldova,
Poland Poland, officially the Republic of Poland, , is a country in Central Europe. Poland is divided into Voivodeships of Poland, sixteen voivodeships and is the fifth most populous member state of the European Union (EU), with over 38 mill ...
, Romania,
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the ...
, Slovenia, Russia (Rostov Oblast, Stavropol Krai, Krasnodar Krai, Moscow Region - non-indigenous distributionEgorov R. (2014). "The first record of ''Cepaea vindobonensis'' (Pfeiffer, 1828) (Stylommatophora: Helicidae) in the central part of European Russia". ''
Malacologica Bohemoslovaca ''Malacologica Bohemoslovaca'' is a peer-reviewed open access scientific journal In academic publishing, a scientific journal is a periodical publication intended to further the progress of science, usually by reporting new research. Cont ...
'' 13: 110-113
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) and
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian invas ...
. In 2021, introduced populations in the United States (New York State) and Canada (Quebec) were reported.Hausdorf, B., Parr, M., Shappell, L. J., Oldeland, J., Robinson, D. G. (2021) The introduction of the European ''Caucasotachea vindobonensis'' (Gastropoda: Helicidae) in North America, its origin and its potential range. ''
Biological Invasions Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary i ...
'
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The original habitat of ''C. vindobonensis'' was most probably open forests in the Balkans, from where it spread over large parts of Central and Eastern Europe after the last glaciation. Nevertheless, it also dwells in several grassland habitats like meadows, steppe and ruderal areas, especially in the north of its distribution area. The highest vertical occurrence is about 1600 m asl in Southern Bulgaria.


Life cycle

All the following information originates from a Greek study. Because Greece is in the southernmost part of the distribution area, the phenology for this species might be quite different in the more northerly countries where it is found. According to the Greek study, ''C. vindobonensis'' becomes mature after its second year, and can reach a maximum life span of about 7 years. The reproductive period in Greece lasts from April to June, with a maximum in May. About 29-67 eggs are laid, having a diameter of about 3 mm, and the juveniles hatch out after 18 days. During hot summer days, these snails rest attached to leaves or the stems of tall plants. Hibernation commences at the end of October to the beginning of November, and the snails come out of hibernation in March.


References


External links

{{Taxonbar, from=Q898580 Helicidae Gastropods described in 1828