Celtis Paniculata
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''Celtis paniculata'', commonly known as tripewood, silky keltis, silky celtis, native hackberry, native celtis, Investigator tree or whitewood, is a
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
tree in the family
Cannabaceae Cannabaceae is a small family of flowering plants, known as the hemp family. As now circumscribed, the family includes about 170 species grouped in about 11 genera, including '' Cannabis'' (hemp), '' Humulus'' (hops) and '' Celtis'' (hackberri ...
native to parts of
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms. It is a phytogeographical floristic region in the Paleotropical kingdom. It was first recognized as a distinct region ...
,
Melanesia Melanesia (, ) is a subregion of Oceania in the southwestern Pacific Ocean. It extends from New Guinea in the west to the Fiji Islands in the east, and includes the Arafura Sea. The region includes the four independent countries of Fiji, Vanu ...
and
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
.


Description

The tree grows to tall and trunk width, though usually much smaller. The trunk is mostly cylindrical and somewhat buttressed at the base. The bark is thin, greyish brown and smooth, with pustular vertical lines.


Leaves

Leaves are elliptical in shape with a prominent leaf tip. Leaves are alternate and simple, smooth, not toothed, long with a pointed tip. The leaf stalk is somewhat channelled and flattened on the top, and in length. Both sides of the leaf are of a dull green shade. With a hand lens, translucent glands may be seen in the leaves. The base of the leaf is often oblique, that is of unequal length on either side of the leaf stem. Leaf venation is more evident on the underside of the leaf. An identifying feature is the intramarginal leaf vein, which starts at the leaf base and travels on either edge of the leaf, a distance of quarter to half the length of the leaf.


Flowers and fruit

Green flowers in axilliary
cymes In botany, an inflorescence is a group or cluster of flowers arranged on a plant's stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a main axis ( ped ...
or
panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...
s appear from December to January. The fruit is a blue to black, round to oval shaped
drupe In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...
, long. containing a single seed which has veiny ridges. Fruit matures from July to March.


Germination and seedlings

Germination takes between 48 and 100 days. The
cotyledon A cotyledon ( ; ; "a cavity, small cup, any cup-shaped hollow", gen. (), ) is a "seed leaf" – a significant part of the embryo within the seed of a plant – and is formally defined as "the embryonic leaf in seed-bearing plants, one or mor ...
s are large, about long by wide, with an apex divided into two acute lobes. The first pairs of juvenile leaves are often toothed. At the tenth leaf stage leaves become oval in shape with a prominent tip.


Taxonomy & naming

This plant first appeared in scientific literature as ''Solenostigma paniculatum'' in 1833, published in the ''Prodromus Florae Norfolkicae'' by the Austrian botanist
Stephan Endlicher Stephan Friedrich Ladislaus Endlicher, also known as Endlicher István László (24 June 1804 – 28 March 1849), was an Austrian Empire, Austrian botanist, numismatist and Sinologist. He was a director of the Botanical Garden of Vienna. Biog ...
. The
type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...
was collected at
Norfolk Island Norfolk Island ( , ; ) is an States and territories of Australia, external territory of Australia located in the Pacific Ocean between New Zealand and New Caledonia, directly east of Australia's Evans Head, New South Wales, Evans Head and a ...
by the artist
Ferdinand Bauer Ferdinand Lucas Bauer (20 January 1760 – 17 March 1826) was an Austrian botanical illustrator who travelled on Matthew Flinders' expedition to Australia. Biography Early life and career Bauer was born in Feldsberg in 1760, the youngest son ...
. The specific epithet ''paniculata'' refers to the tufted and branched flowers. In 1848
Jules Émile Planchon Jules Émile Planchon (21 March 1823 – 1 April 1888) was a French botanist born in Ganges, Hérault. He was a brother of Gustave Planchon who also studied medicinal plants. Biography After receiving his Doctorate of Science at the Universit ...
moved the species to the current genus.


Origin of the name 'Investigator Tree'

In 1802 the explorer
Matthew Flinders Captain (Royal Navy), Captain Matthew Flinders (16 March 1774 – 19 July 1814) was a British Royal Navy officer, navigator and cartographer who led the first littoral zone, inshore circumnavigate, circumnavigation of mainland Australia, then ...
discovered and named Sweers Island in the
Gulf of Carpentaria The Gulf of Carpentaria is a sea off the northern coast of Australia. It is enclosed on three sides by northern Australia and bounded on the north by the eastern Arafura Sea, which separates Australia and New Guinea. The northern boundary ...
during his circumnavigation of Australia in the sloop HMS Investigator, and he carved the word 'Investigator' into a tree on the island, a specimen of ''Celtis paniculata''. The tree became well known amongst later explorers, who also carved the names of their vessels into the tree. When the tree was damaged by a
tropical cyclone A tropical cyclone is a rapidly rotating storm system with a low-pressure area, a closed low-level atmospheric circulation, strong winds, and a spiral arrangement of thunderstorms that produce heavy rain and squalls. Depending on its locat ...
in 1887, part of the trunk was removed to the Queensland Museum.


Distribution and habitat

''C. paniculatta'' has a wide natural distribution in tropical Asia and the Pacific, including
Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...
, the
Moluccas The Maluku Islands ( ; , ) or the Moluccas ( ; ) are an archipelago in the eastern part of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located in West Melanesi ...
, the
Lesser Sunda Islands The Lesser Sunda Islands (, , ), now known as Nusa Tenggara Islands (, or "Southeast Islands"), are an archipelago in the Indonesian archipelago. Most of the Lesser Sunda Islands are located within the Wallacea region, except for the Bali pro ...
, the
Caroline Islands The Caroline Islands (or the Carolines) are a widely scattered archipelago of tiny islands in the western Pacific Ocean, to the north of New Guinea. Politically, they are divided between the Federated States of Micronesia (FSM) in the cen ...
,
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
, the
Solomon Islands Solomon Islands, also known simply as the Solomons,John Prados, ''Islands of Destiny'', Dutton Caliber, 2012, p,20 and passim is an island country consisting of six major islands and over 1000 smaller islands in Melanesia, part of Oceania, t ...
,
Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (; ), is an island country in Melanesia located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of northern Australia, northeast of New Caledonia, east o ...
,
New Caledonia New Caledonia ( ; ) is a group of islands in the southwest Pacific Ocean, southwest of Vanuatu and east of Australia. Located from Metropolitan France, it forms a Overseas France#Sui generis collectivity, ''sui generis'' collectivity of t ...
, and the
Northern Territory The Northern Territory (abbreviated as NT; known formally as the Northern Territory of Australia and informally as the Territory) is an states and territories of Australia, Australian internal territory in the central and central-northern regi ...
and
Queensland Queensland ( , commonly abbreviated as Qld) is a States and territories of Australia, state in northeastern Australia, and is the second-largest and third-most populous state in Australia. It is bordered by the Northern Territory, South Austr ...
. In Australia it occurs from
Kiama Kiama may refer to: * Electoral district of Kiama, a seat in the New South Wales Legislative Assembly * Kiama, New South Wales, a town * Kiama (spider), a genus of spiders *Municipality of Kiama The Municipality of Kiama is a local government ar ...
(34° S) in
New South Wales New South Wales (commonly abbreviated as NSW) is a States and territories of Australia, state on the Eastern states of Australia, east coast of :Australia. It borders Queensland to the north, Victoria (state), Victoria to the south, and South ...
to
Cape York Peninsula The Cape York Peninsula is a peninsula located in Far North Queensland, Australia. It is the largest wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth's last wild places. Mexico City: Agrupación Sierra Madre, ...
, as well as on
Norfolk Island Norfolk Island ( , ; ) is an States and territories of Australia, external territory of Australia located in the Pacific Ocean between New Zealand and New Caledonia, directly east of Australia's Evans Head, New South Wales, Evans Head and a ...
. It will grow in a variety of forest types, including sandy beach forests,
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
s, monsoon forests and
sclerophyll forests Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes (the distance between leaves along the stem) and leaf orientation which is parallel or oblique to direct ...
, and on a variety of soil types, including soils derived from
basalt Basalt (; ) is an aphanite, aphanitic (fine-grained) extrusive igneous rock formed from the rapid cooling of low-viscosity lava rich in magnesium and iron (mafic lava) exposed at or very near the planetary surface, surface of a terrestrial ...
,
granite Granite ( ) is a coarse-grained (phanerite, phaneritic) intrusive rock, intrusive igneous rock composed mostly of quartz, alkali feldspar, and plagioclase. It forms from magma with a high content of silica and alkali metal oxides that slowly coo ...
and
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
. It is often seen in coastal areas, but in northern parts of Queensland it can be found up to an altitude of .


Ecology & uses

''Celtis paniculata'' is a food tree for the blue beak, common aeroplane, tailed emperor and other butterflies. Many rainforest pigeons and other birds feed on the fruit. An endangered population of the beetle Menippus darcyi feeds on ''Celtis paniculata''. The tree produces a useful general purpose timber, with a specific gravity of 0.70.


Gallery

File:Investigator Tree Celtis paniculata Bass Point.jpg , Growing at Bass Point, NSW File:Image_of_the_Investigator_tree_on_Sweers_Island_in_1871.tiff , The original " Investigator Tree" on Sweers Island in 1871 File:Celtis paniculata seedling.jpg ,
Cotyledon A cotyledon ( ; ; "a cavity, small cup, any cup-shaped hollow", gen. (), ) is a "seed leaf" – a significant part of the embryo within the seed of a plant – and is formally defined as "the embryonic leaf in seed-bearing plants, one or mor ...
s File:Celtis paniculata leaf scan.jpg , Mature leaves


References

{{Taxonbar, from=Q2682213 paniculata Rosales of Australia Trees of Australia Flora of Queensland Flora of New South Wales Flora of Norfolk Island Flora of Malesia Flora of the Caroline Islands Flora of New Guinea Flora of the Solomon Islands (archipelago) Flora of Vanuatu Flora of New Caledonia Taxa named by Stephan Endlicher Plants described in 1833