The copper shark (''Carcharhinus brachyurus''), bronze whaler, or narrowtooth shark is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

requiem shark Requiem sharks are sharks of the family Carcharhinidae in the order Carcharhiniformes. They are migratory, live-bearing sharks of warm seas (sometimes of brackish or fresh water) and include such species as the bull shark, lemon shark, blac ...

found mostly in

temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...

latitude In geography, latitude is a geographic coordinate system, geographic coordinate that specifies the north-south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from −90° at t ...

s. It is distributed in a number of separate populations in the northeastern and southwestern

Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...

, off

southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...

, in the northwestern and eastern

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is bounded by the cont ...

, and around

Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...

and

New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...

, with scattered reports from

equator The equator is the circle of latitude that divides Earth into the Northern Hemisphere, Northern and Southern Hemisphere, Southern Hemispheres of Earth, hemispheres. It is an imaginary line located at 0 degrees latitude, about in circumferen ...

ial regions. The species can be found from

brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...

rivers and

estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...

to shallow bays and harbors, to offshore waters deep or more. Females are found apart from males for most of the year, and conduct seasonal migrations. A large species reaching long, the copper shark is difficult to distinguish from other large requiem sharks. It is characterized by its narrow, hook-shaped upper teeth, lack of a prominent ridge between the

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

s, and plain bronze coloration. Feeding mainly on

cephalopod A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symm ...

bony fish Osteichthyes ( ; ), also known as osteichthyans or commonly referred to as the bony fish, is a Biodiversity, diverse clade of vertebrate animals that have endoskeletons primarily composed of bone tissue. They can be contrasted with the Chondricht ...

es, and other

cartilaginous fish Chondrichthyes (; ) is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fish'', which have skeleto ...

es, the copper shark is a fast-swimming

predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...

that has been known to hunt in large groups, using their numbers to their advantage; however, for most of the time they remain solitary. Off

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...

, this species associates closely with the annual

sardine run The KwaZulu-Natal sardine run of southern Africa occurs from May through July when billions of sardines – or more specifically the Southern African pilchard ''Sardinops sagax'' – spawn in the cool waters of the Agulhas Bank and move northwar ...

, involving millions of

southern African pilchard ''Sardinops'' is a monotypic genus of sardines of the family Alosidae. The only member of the genus is ''Sardinops sagax''. It is found in the Indo-Pacific and East Pacific oceans. Its length is up to . It has numerous common or vernacular names, ...

(''Sardinops sagax''). Like other requiem sharks, it is

viviparous In animals, viviparity is development of the embryo inside the body of the mother, with the maternal circulation providing for the metabolic needs of the embryo's development, until the mother gives birth to a fully or partially developed juve ...

, with the developing

embryo An embryo ( ) is the initial stage of development for a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sp ...

s mainly nourished through a

placenta The placenta (: placentas or placentae) is a temporary embryonic and later fetal organ that begins developing from the blastocyst shortly after implantation. It plays critical roles in facilitating nutrient, gas, and waste exchange between ...

l connection formed from the depleted

yolk sac The yolk sac is a membranous wikt:sac, sac attached to an embryo, formed by cells of the hypoblast layer of the bilaminar embryonic disc. This is alternatively called the umbilical vesicle by the Terminologia Embryologica (TE), though ''yolk sac' ...

. Females bear litters of 7 to 24 pups every other year in coastal nursery areas, after a

gestation period In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once i ...

of 12 or perhaps as long as 21 months. It is extremely slow-growing, with males and females not reaching maturity until 13–19 and 19–20 years of age respectively. This species is valued by

commercial Commercial may refer to: * (adjective for) commerce, a system of voluntary exchange of products and services ** (adjective for) trade, the trading of something of economic value such as goods, services, information or money * a dose of advertising ...

and recreational fisheries throughout its range, and utilized as food. The species population size is unknown, but the IUCN's Red List assesses the species as vulnerable because it is very susceptible to population depletion due to its low growth and reproductive rates and because its numbers are believed to have declined in some areas. Copper sharks only attack humans infrequently, but the species places tenth in the number of unprovoked attacks on people. However, the actual number of recorded shark attacks is low – 15 non-fatal, unprovoked attacks and only one fatal unprovoked attack.

Taxonomy

Because of its very patchy range, the copper shark has been scientifically described several times from different locales. The earliest valid description is presently considered to be British

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

Albert Günther Albert Karl Ludwig Gotthilf Günther , also Albert Charles Lewis Gotthilf Günther (3October 18301February 1914), was a German-born British zoologist, ichthyologist, and herpetologist. Günther is ranked the second-most productive reptile tax ...

's account of ''Carcharias brachyurus'' in the 1870 eighth volume of ''Catalogue of the fishes in the British Museum''. The earliest name was once thought to be

Auguste Duméril Auguste Henri André Duméril (30 November 1812 – 12 November 1870) was a French zoologist. His father, André Marie Constant Duméril (1774–1860), was also a zoologist. In 1869 he was elected as a member of the Académie des sciences. Dum ...

's 1865 ''Carcharias remotus'', until it was found that the

type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...

associated with that name is actually a

blacknose shark The blacknose shark (''Carcharhinus acronotus'') is a species of requiem shark, belonging to the family Carcharhinidae, common in the tropical and subtropical waters of the western Atlantic Ocean. This species generally inhabits coastal seagrass, ...

(''C. acronotus''). Thus, this shark was often referred to as ''C. remotus'' in older literature. An even earlier name,

Richard Owen Sir Richard Owen (20 July 1804 – 18 December 1892) was an English biologist, comparative anatomy, comparative anatomist and paleontology, palaeontologist. Owen is generally considered to have been an outstanding naturalist with a remarkabl ...

's 1853 ''Galeolamna greyi'', is of questionable taxonomic status as it was based solely on a set of now-destroyed jaws that may or may not have belonged to a copper shark. Modern authors have assigned this species to the genus ''Carcharhinus''. The

specific epithet In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...

''brachyurus'' is derived from the

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

''brachys'' ("short") and ''oura'' ("tail"). The name "whaler" originated in the 19th century, applied by the crews of whaling vessels in the Pacific who saw large sharks of various species congregating around harpooned whale carcasses. This species may also be referred to as black-tipped whaler, cocktail shark or cocktail whaler, or New Zealand whaler, as well as by the shortened "bronze", "bronzie", or "cocktail". Günther originally referred to four

syntype In biological nomenclature, a syntype is any one of two or more biological types that is listed in a description of a taxon where no holotype was designated. Precise definitions of this and related terms for types have been established as part o ...

s: a stuffed specimen from

Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...

and another from

, which have since been lost, and two

fetus A fetus or foetus (; : fetuses, foetuses, rarely feti or foeti) is the unborn offspring of a viviparous animal that develops from an embryo. Following the embryonic development, embryonic stage, the fetal stage of development takes place. Pren ...

es from

that were later discovered to be

bull shark The bull shark (''Carcharhinus leucas''), also known as the Zambezi shark (informally zambi) in Africa and Lake Nicaragua shark in Nicaragua, is a species of requiem shark commonly found worldwide in warm, shallow waters along coasts and in riv ...

s (''C. leucas''). In the interests of taxonomic stability, in 1982

Jack Garrick John Andrew Frank Garrick (24 March 1928 – 30 August 2018) was a New Zealand ichthyologist. Career Garrick specialised in elasmobranchs and published many books and articles about shark and ray biology. In 1982, he published a thorough taxonom ...

designated a long female caught off

Whanganui Whanganui, also spelt Wanganui, is a city in the Manawatū-Whanganui region of New Zealand. The city is located on the west coast of the North Island at the mouth of the Whanganui River, New Zealand's longest navigable waterway. Whanganui is ...

, New Zealand, as a new type specimen.

Phylogeny and evolution

The first efforts to determine the evolutionary relationships of the copper shark were based on

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...

and returned inconclusive results: in 1982 Jack Garrick placed it by itself as a grouping within ''Carcharhinus'', while in 1988

Leonard Compagno Leonard Joseph Victor Compagno (1943-2024) was an international authority on shark taxonomy and the author of many scientific papers and books on the subject, best known of which is his 1984 catalogue of shark species produced for the Food and Agri ...

placed it in an informal "transitional group" that also contained the

(''C. acronotus''), the

blacktip reef shark The blacktip reef shark (''Carcharhinus melanopterus'') is a species of requiem shark, in the family (biology), family Carcharhinidae, which can be easily identified by the prominent black tips on its fins (especially on the first dorsal fin and ...

(''C. melanopterus''), the nervous shark (''C. cautus''), the

silky shark The silky shark (''Carcharhinus falciformis''), also known by numerous names such as blackspot shark, gray whaler shark, olive shark, ridgeback shark, sickle shark, sickle-shaped shark and sickle silk shark, is a species of requiem shark, in the ...

(''C. falciformis''), and the

night shark The night shark (''Carcharhinus signatus'') is a species of requiem shark, in the family Carcharhinidae, found in the temperate and tropical waters of the Atlantic Ocean. An inhabitant of the outer continental shelf and upper continental slope, ...

(''C. signatus''). Gavin Naylor's 1992

allozyme Alloenzymes (or also called allozymes) are variant forms of an enzyme which differ structurally but not functionally from other allozymes coded for by different alleles at the same locus. These are opposed to isozymes, which are enzymes that p ...

study concluded that the closest relative of the copper shark is the

spinner shark The spinner shark (''Carcharhinus brevipinna'') is a type of requiem shark, in the family Carcharhinidae, named for the spinning leaps it makes as a part of its feeding strategy. This species occurs in tropical and warm temperate waters worldwid ...

(''C. brevipinna''), but could not resolve their wider relationships with the rest of the genus.

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

ized teeth from the copper shark have been recovered from the Chilcatay Formation in

Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...

dating to the

Early Miocene The Early Miocene (also known as Lower Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages: the Aquitanian age, Aquitanian and Burdigalian stages. The sub-epoch lasted from 23.03 ± 0.05 annum, Ma to ...

Pungo River The Pungo River is a river in eastern North Carolina, United States. It originally began in the Great Dismal Swamp in Washington County, North Carolina; the upper part of the river has since been supplanted by the Pungo River Canal, dug in the 1 ...

North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...

, dating to the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

(23–5.3 Ma), from

Tuscany Tuscany ( ; ) is a Regions of Italy, region in central Italy with an area of about and a population of 3,660,834 inhabitants as of 2025. The capital city is Florence. Tuscany is known for its landscapes, history, artistic legacy, and its in ...

, dating to the

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58 and from

Costa Mesa Costa may refer to: Biology * Rib (Latin: ''costa''), in vertebrate anatomy * Costa (botany), the central strand of a plant leaf or thallus * Costa (coral), a stony rib, part of the skeleton of a coral * Costa (entomology), the leading edge o ...

California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...

, dating to the

Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...

(126,000–12,000 years ago).

Distribution and habitat

The copper shark is the only member of its genus largely found in

rather than

tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...

waters, in temperatures above . It is widely distributed but as disjunct regional populations with little to no interchange between them. In the Atlantic, this shark occurs from the

Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...

Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...

and the

Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...

, off

Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...

, and off

Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...

and

(where there may be two separate populations), with infrequent records from

Mauritania Mauritania, officially the Islamic Republic of Mauritania, is a sovereign country in Maghreb, Northwest Africa. It is bordered by the Atlantic Ocean to the west, Western Sahara to Mauritania–Western Sahara border, the north and northwest, ...

, the

Gulf of Guinea The Gulf of Guinea (French language, French: ''Golfe de Guinée''; Spanish language, Spanish: ''Golfo de Guinea''; Portuguese language, Portuguese: ''Golfo da Guiné'') is the northeasternmost part of the tropical Atlantic Ocean from Cape Lopez i ...

, and possibly the

Gulf of Mexico The Gulf of Mexico () is an oceanic basin and a marginal sea of the Atlantic Ocean, mostly surrounded by the North American continent. It is bounded on the northeast, north, and northwest by the Gulf Coast of the United States; on the southw ...

. In the

Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...

, it is found from the

East China Sea The East China Sea is a marginal sea of the Western Pacific Ocean, located directly offshore from East China. China names the body of water along its eastern coast as "East Sea" (, ) due to direction, the name of "East China Sea" is otherwise ...

Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...

(excluding

Hokkaido is the list of islands of Japan by area, second-largest island of Japan and comprises the largest and northernmost prefectures of Japan, prefecture, making up its own list of regions of Japan, region. The Tsugaru Strait separates Hokkaidō fr ...

) and southern

Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...

, off southern

(mostly between

Sydney Sydney is the capital city of the States and territories of Australia, state of New South Wales and the List of cities in Australia by population, most populous city in Australia. Located on Australia's east coast, the metropolis surrounds Syd ...

and

Perth Perth () is the list of Australian capital cities, capital city of Western Australia. It is the list of cities in Australia by population, fourth-most-populous city in Australia, with a population of over 2.3 million within Greater Perth . The ...

but occasionally further north), and around

but not as far as the

Kermadec Islands The Kermadec Islands ( ; ) are a subtropical island arc in the South Pacific Ocean northeast of New Zealand's North Island, and a similar distance southwest of Tonga. The islands are part of New Zealand. They are in total area and uninhabit ...

; there are also unconfirmed reports from the

Seychelles Seychelles (, ; ), officially the Republic of Seychelles (; Seychellois Creole: ), is an island country and archipelagic state consisting of 155 islands (as per the Constitution) in the Indian Ocean. Its capital and largest city, Victoria, ...

and the

Gulf of Thailand The Gulf of Thailand (), historically known as the Gulf of Siam (), is a shallow inlet adjacent to the southwestern South China Sea, bounded between the southwestern shores of the Indochinese Peninsula and the northern half of the Malay Peninsula. ...

. In the eastern

, it occurs from northern

Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...

, and from

Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...

Point Conception Point Conception (Chumashan languages, Chumash: ''Humqaq'') is a headland along the Gaviota Coast in southwestern Santa Barbara County, California, United States. It is the point where the Santa Barbara Channel meets the Pacific Ocean, and as ...

, including the

Gulf of California The Gulf of California (), also known as the Sea of Cortés (''Mar de Cortés'') or Sea of Cortez, or less commonly as the Vermilion Sea (''Mar Vermejo''), is a marginal sea of the Pacific Ocean that separates the Baja California peninsula from ...

. The copper shark is common off parts of Argentina, South Africa, Australia, and New Zealand, and rare elsewhere; in many areas its range is ill-defined because of confusion with other species. The shark is one of the most commonly found in the waters of New Zealand. Copper sharks can be found from the

surf zone The surf zone or breaker zone is the nearshore part of a body of open water between the line at which the waves break and the shore. As ocean surface waves approach a shore, they interact with the bottom, wave shoaling, get taller and steeper, an ...

to slightly beyond the

continental shelf A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an islan ...

in the

open ocean The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the surfa ...

, diving to depths of or more. This species commonly enters very shallow

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

s, including

bay A bay is a recessed, coastal body of water that directly connects to a larger main body of water, such as an ocean, a lake, or another bay. A large bay is usually called a ''gulf'', ''sea'', ''sound'', or ''bight''. A ''cove'' is a small, ci ...

shoal In oceanography, geomorphology, and Earth science, geoscience, a shoal is a natural submerged ridge, bank (geography), bank, or bar that consists of, or is covered by, sand or other unconsolidated material, and rises from the bed of a body ...

s, and

harbor A harbor (American English), or harbour (Commonwealth English; see spelling differences), is a sheltered body of water where ships, boats, and barges can be moored. The term ''harbor'' is often used interchangeably with ''port'', which is ...

s, and also inhabits rocky areas and offshore islands. It is tolerant of low and changing

salinities Salinity () is the saltiness or amount of salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensionless and equal to ...

, and has been reported from

and the lower reaches of large

river A river is a natural stream of fresh water that flows on land or inside Subterranean river, caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of ...

s. Juveniles inhabit inshore waters less than deep throughout the year, while adults tend to be found further offshore and regularly approach the coast only in spring and summer, when large aggregations can be readily observed in shallow water. Populations of copper sharks in both hemispheres perform seasonal migrations, in response to temperature changes, reproductive events, and/or prey availability; the movement patterns differ with sex and age. Adult females and juveniles spend winter in the subtropics and generally shift to higher latitudes as spring nears, with pregnant females also moving towards the coast to give birth in inshore nursery areas. Adult males remain in the

subtropics The subtropical zones or subtropics are geographical and climate zones immediately to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately ...

for most of the year, except in late winter or spring when they also move into higher latitudes, in time to encounter and mate with post-partum females dispersing from the nurseries. During migrations, individual sharks have been recorded traveling up to . It is philopatric, returning to the same areas year after year.

Description

The copper shark has a slender, streamlined body with a slightly arched profile just behind the head. The snout is rather long and pointed, with the nostrils preceded by low flaps of skin. The round, moderately large eyes are equipped with

nictitating membrane The nictitating membrane (from Latin '' nictare'', to blink) is a transparent or translucent third eyelid present in some animals that can be drawn across the eye from the medial canthus to protect and moisten it while maintaining vision. Most ...

s (protective third eyelids). The mouth has short, subtle furrows at the corners and contains 29–35 upper tooth rows and 29–33 lower tooth rows. The teeth are serrated with single narrow cusps; the upper teeth have a distinctive hooked shape and become more angled towards the corners of the jaw, while the lower teeth are upright. The upper teeth of adult males are longer, narrower, more curved, and more finely serrated than those of adult females and juveniles. The five pairs of

gill slit Gill slits are individual openings to gills, i.e., multiple gill arches, which lack a single outer cover. Such gills are characteristic of cartilaginous fish such as sharks and rays, as well as deep-branching vertebrates such as lampreys. In c ...

s are fairly long. The

pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...

s are large, pointed, and falcate (sickle-shaped). The first

is tall, with a pointed apex and a concave trailing margin; its origin lies about even with the tips of the pectoral fins. The second dorsal fin is small and low, and positioned about opposite to the

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

. There is usually no ridge between the dorsal fins. The

caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...

has a well-developed lower lobe and a deep ventral notch near the tip of the upper lobe. This species is bronze to olive-gray above with a metallic sheen and sometimes a pink cast, darkening towards the fin tips and margins but not conspicuously so; the color fades quickly to a dull gray-brown after death. The underside is white, which extends onto the flanks as a prominent band. The copper shark is easily mistaken for other large ''Carcharhinus'' species, particularly the

dusky shark The dusky shark (''Carcharhinus obscurus'') is a species of requiem shark, in the family Carcharhinidae, occurring in tropical and warm-temperate continental seas worldwide. A generalist apex predator, the dusky shark can be found from the coas ...

(''C. obscurus''), but can be identified by its upper tooth shape, absent or weak interdorsal ridge, and lack of obvious fin markings. It reportedly reaches a maximum length of and weight of . Carcharhinus brachyurus phuket.JPG, The copper shark can be difficult to distinguish from other large ''Carcharhinus'' species Carcharhinus brachyurus jaws.jpg, Jaws Carcharhinus brachyurus upper teeth.jpg, Upper teeth Carcharhinus brachyurus lower teeth.jpg, Lower teeth

Biology and ecology

Fast but active, the copper shark may be encountered alone, in pairs, or in loosely organized

school A school is the educational institution (and, in the case of in-person learning, the Educational architecture, building) designed to provide learning environments for the teaching of students, usually under the direction of teachers. Most co ...

s containing up to hundreds of individuals. Some aggregations seem to form for reproductive purposes, while others form to concentrations of food. This species may fall prey to larger sharks and killer whales. Known

parasite Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted str ...

s of the copper shark include the

tapeworm Eucestoda, commonly referred to as tapeworms, is the larger of the two subclasses of flatworms in the class Cestoda (the other subclass being Cestodaria). Larvae have six posterior hooks on the scolex (head), in contrast to the ten-hooked Ce ...

s ''Cathetocephalus australis'', ''Dasyrhynchus pacificus'' and ''D. talismani'', ''Floriceps minacanthus'', ''Phoreiobothrium robertsoni'', and ''Pseudogrillotia spratti'', the

leech Leeches are segmented parasitism, parasitic or Predation, predatory worms that comprise the Class (biology), subclass Hirudinea within the phylum Annelida. They are closely related to the Oligochaeta, oligochaetes, which include the earthwor ...

''Stibarobdella macrothela'', and the fluke ''Otodistomum veliporum''.

Feeding

The copper shark feeds more towards the bottom of the

water column The (oceanic) water column is a concept used in oceanography to describe the physical (temperature, salinity, light penetration) and chemical ( pH, dissolved oxygen, nutrient salts) characteristics of seawater at different depths for a defined ...

than the top, consuming

s, including

squid A squid (: squid) is a mollusc with an elongated soft body, large eyes, eight cephalopod limb, arms, and two tentacles in the orders Myopsida, Oegopsida, and Bathyteuthida (though many other molluscs within the broader Neocoleoidea are also ...

(''

Loligo ''Loligo'' is a genus of squid and one of the most representative and widely distributed groups of myopsid squid. The genus was first described by Jean Baptiste Lamarck in 1798. However, the name had been used earlier than Lamarck ( Schneider, ...

'' spp.),

cuttlefish Cuttlefish, or cuttles, are Marine (ocean), marine Mollusca, molluscs of the order (biology), suborder Sepiina. They belong to the class (biology), class Cephalopoda which also includes squid, octopuses, and nautiluses. Cuttlefish have a unique ...

es, and

octopus An octopus (: octopuses or octopodes) is a soft-bodied, eight-limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttlefish, and nautiloids. Like oth ...

;

es, including gurnards,

flatfish A flatfish is a member of the Ray-finned fish, ray-finned demersal fish Order (biology), suborder Pleuronectoidei, also called the Heterosomata. In many species, both eyes lie on one side of the head, one or the other migrating through or around ...

es, hakes,

catfish Catfish (or catfishes; order (biology), order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Catfish are common name, named for their prominent barbel (anatomy), barbels, which resemble a cat's whiskers, though not ...

es, jacks, Australian salmon,

Atlantic salmon The Atlantic salmon (''Salmo salar'') is a species of ray-finned fish in the family Salmonidae. It is the third largest of the Salmonidae, behind Hucho taimen, Siberian taimen and Pacific Chinook salmon, growing up to a meter in length. Atlan ...

, mullets,

sea bream Sparidae is a family of ray-finned fishes belonging to the order Spariformes, the seabreams and porgies, although they were traditionally classified in the order Perciformes. The over 150 species are found in shallow and deep marine waters in te ...

s, smelts,

tuna A tuna (: tunas or tuna) is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae ( mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bul ...

sardine Sardine and pilchard are common names for various species of small, oily forage fish in the herring suborder Clupeoidei. The term "sardine" was first used in English during the early 15th century; a somewhat dubious etymology says it com ...

Cape horse mackerel The Cape horse mackerel (''Trachurus capensis'') is a mackerel-like species in the family Carangidae. It is a pelagic species of the south eastern Atlantic Ocean which is a target of fisheries, mainly as bycatch. Description The Cape horse macke ...

, and

anchovies An anchovy is a small, common forage fish of the family Engraulidae. Most species are found in marine waters, but several will enter brackish water, and some in South America are restricted to fresh water. More than 140 species are placed in 1 ...

; and

es, including dogfish sharks (''Squalus'' spp.),

stingray Stingrays are a group of sea Batoidea, rays, a type of cartilaginous fish. They are classified in the suborder Myliobatoidei of the order Myliobatiformes and consist of eight families: Hexatrygonidae (sixgill stingray), Plesiobatidae (deepwate ...

skate Skate or Skates may refer to: Fish *Skate (fish), several genera of fish belonging to the family Rajidae * Pygmy skates, several genera of fish belonging to the family Gurgesiellidae * Smooth skates or leg skates, several genera of fish belongin ...

electric ray The electric rays are a group of rays, flattened cartilaginous fish with enlarged pectoral fins, composing the order Torpediniformes . They are known for being capable of producing an electric discharge, ranging from 8 to 220 volts, depending ...

s, and

sawfish Sawfish, also known as carpenter sharks, are a family of very large rays characterized by a long, narrow, flattened rostrum, or nose extension, lined with sharp transverse teeth, arranged in a way that resembles a saw. They are among the lar ...

es. Cephalopods and cartilaginous fishes become relatively more important food for sharks over long. Young sharks also consume scyphozoan jellyfish and

crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...

s, including mud shrimps (''

Callianassa ''Callianassa'' is a genus of mud shrimps, in the family Callianassidae. Three of the species in this genus ('' C. candida'', '' C. tyrrhena'' and '' C. whitei'') have been split off into a new genus, '' Pestarella'', while others such as '' Ca ...

'') and

penaeid Penaeidae is a family of marine crustaceans in the suborder Dendrobranchiata, which are often referred to as penaeid shrimp or penaeid prawns. The Penaeidae contain many species of economic importance, such as the tiger prawn, whiteleg shrimp, ...

prawn Prawn is a common name for small aquatic crustaceans with an exoskeleton An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the intern ...

s. It does not attack

marine mammals Marine mammals are mammals that rely on marine ecosystems for their existence. They include animals such as cetaceans, pinnipeds, sirenians, sea otters and polar bears. They are an informal group, unified only by their reliance on marine env ...

, though it has been known to scavenge (rarely) on

dolphin A dolphin is an aquatic mammal in the cetacean clade Odontoceti (toothed whale). Dolphins belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontopori ...

s that had succumbed to fishing nets. The predominant prey of this species off South Africa is the

(''Sardinops sagax''), which comprise 69–95% of its diet. Every winter, schools of copper sharks follow the "run" of the pilchard from the Eastern Cape to

KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN) is a Provinces of South Africa, province of South Africa that was created in 1994 when the government merged the Zulu people, Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu language, Zulu) and ...

. The gathering of millions of

forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish that feed on planktons (i.e. planktivores) and other small aquatic organisms (e.g. krill). They are in turn preyed upon by various predators including larger fish, seabirds ...

attract a multitude of predators, including several species of sharks, of which copper sharks are the most numerous. Large numbers of copper sharks have been observed hunting together in a seemingly cooperative fashion. Small schooling fish are "herded" into a tight ball, whereupon each shark swims through in turn with its mouth open to feed. For groups of tuna and larger prey, the pursuing sharks may adopt a "wing" formation to force their quarry closer together, with each shark targeting a particular fish and attacking in turn. In

False Bay False Bay (Afrikaans: ''Valsbaai'') is a body of water in the Atlantic Ocean between the mountainous Cape Peninsula and the Hottentots Holland Mountains in the extreme south-west of South Africa. The mouth of the bay faces south and is demarc ...

, South Africa, this species reportedly follows

seine net Seine fishing (or seine-haul fishing; ) is a method of fishing that employs a surrounding net, called a seine, that hangs vertically in the water with its bottom edge held down by weights and its top edge buoyed by floats. Seine nets can be de ...

fishing vessels.

Life history

Like other members of its family, the copper shark is

, in which the

of the developing

, once depleted, is converted into a

l connection through which the mother delivers nourishment. Adult females have one functional

ovary The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...

, on the right, and two functional

uterus The uterus (from Latin ''uterus'', : uteri or uteruses) or womb () is the hollow organ, organ in the reproductive system of most female mammals, including humans, that accommodates the embryonic development, embryonic and prenatal development, f ...

es. The male bites the female as a prelude to mating. In the Southern Hemisphere, mating takes place from October to December (spring and early summer), when both sexes have migrated into offshore waters at higher latitudes. Birthing seems to occur from June to January, peaking in October and November. Female copper sharks make use of shallow inshore habitats, varying from stretches of open coast to more sheltered bays and inlets, as nurseries. These nurseries provide abundant food and reduce the likelihood of predation by larger members of the species. Known and suspected nursery areas occur off northern

North Island The North Island ( , 'the fish of Māui', historically New Ulster) is one of the two main islands of New Zealand, islands of New Zealand, separated from the larger but less populous South Island by Cook Strait. With an area of , it is the List ...

from Waimea Inlet to

Hawke Bay Hawke Bay (), formerly named ''Hawke's Bay'', is a large bay on the east coast of the North Island of New Zealand, surrounded by the Hawke's Bay region. It stretches from Māhia Peninsula in the northeast to Cape Kidnappers / Te Kauwae-a-Mā ...

for New Zealand sharks, off Albany, in and around

Gulf St Vincent Gulf St Vincent, sometimes referred to as St Vincent Gulf, St Vincent's Gulf or Gulf of St Vincent, is the eastern of two large inlets of water on the southern coast of Australia, in the state of South Australia, the other being the larger Sp ...

, and in

Port Phillip Bay Port Phillip ( Kulin: ''Narm-Narm'') or Port Phillip Bay is a horsehead-shaped enclosed bay on the central coast of southern Victoria, Australia. The bay opens into the Bass Strait via a short, narrow channel known as The Rip, and is comple ...

for Australian sharks, off Niigata (Japan) for northwestern Pacific sharks, off the

Eastern Cape The Eastern Cape ( ; ) is one of the nine provinces of South Africa. Its capital is Bhisho, and its largest city is Gqeberha (Port Elizabeth). Due to its climate and nineteenth-century towns, it is a common location for tourists. It is also kno ...

for South Africa sharks, off

Rhodes Rhodes (; ) is the largest of the Dodecanese islands of Greece and is their historical capital; it is the List of islands in the Mediterranean#By area, ninth largest island in the Mediterranean Sea. Administratively, the island forms a separ ...

(Greece),

Nice Nice ( ; ) is a city in and the prefecture of the Alpes-Maritimes department in France. The Nice agglomeration extends far beyond the administrative city limits, with a population of nearly one million Al Hoceima Al Hoceima () is a city in the north of Morocco, on the northern edge of the Rif Mountains and on the Mediterranean coast. It is the capital city of the Al Hoceima Province. The city is a known tourist destination despite its small size. It has ...

(Morocco) for Mediterranean sharks, off

Río de Oro Río de Oro (, Spanish for "River of Gold"; , , often transliterated as ''Oued Edhahab'') is the southern geographic region of Western Sahara. It was, with Saguia el-Hamra, one of the two territories that formed the Spanish province of S ...

(Western Sahara) for northwest African sharks, off

Rio de Janeiro Rio de Janeiro, or simply Rio, is the capital of the Rio de Janeiro (state), state of Rio de Janeiro. It is the List of cities in Brazil by population, second-most-populous city in Brazil (after São Paulo) and the Largest cities in the America ...

(Brazil) and

Buenos Aires Buenos Aires, controlled by the government of the Autonomous City of Buenos Aires, is the Capital city, capital and largest city of Argentina. It is located on the southwest of the Río de la Plata. Buenos Aires is classified as an Alpha− glob ...

and

Bahía Blanca Bahía Blanca (; English: ''White Bay''), colloquially referred to by its own local inhabitants as simply Bahía, is a city in the Buenos Aires Province, Buenos Aires province of Argentina, centered on the northwestern end of the eponymous Blanc ...

(Argentina) for southwestern Atlantic sharks, and off

Paita Paita is a city in northwestern Peru. It is the capital of the Paita Province which is in the Piura Region. It is a leading seaport in the region. Paita is located 1,089 km northwest of the country's capital Lima, and 57 km northwest of ...

and Guanape Cove (Peru), in

Sebastián Vizcaíno Bay Sebastián Vizcaíno Bay (Spanish: ''Bahía de Sebastián Vizcaíno'') is a prominent bay along the west coast of the Baja California Peninsula in northwestern Mexico. It is named after explorer Sebastián Vizcaíno. Geography The bay lies on ...

(Mexico), and in and around

San Diego Bay San Diego Bay is a natural harbor and deepwater port in San Diego County, California, near the Mexico–United States border. The bay, which is long and wide, is the third largest of the three large, protected natural bays on California's of ...

for eastern Pacific sharks. Most sources estimate a

of 12 months, though some data support the interpretation of a 15–21 month long gestation period instead. Females produce litters every other year, with the number of pups ranging from 7 to 24 and averaging 15 or 16. Females off California and the Baja Peninsula tend to bear fewer young relative to other parts of the world. The newborns measure long. The copper shark is among the slowest-growing ''Carcharhinus'' species: off South Africa, males reach

sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. ''Puberty'' is the biological process of sexual maturation, while ''adulthood'', the condition of being socially recognized ...

at long and an age of 13–19 years, while females mature at long and an age of 19–20 years. Females off Australia mature at a comparable length of , while females off Argentina mature at a slightly smaller length of . The

maximum lifespan Maximum life span (or, for humans, maximum reported age at death) is a measure of the maximum amount of time one or more members of a population have been observed to survive between birth and death. The term can also denote an estimate of the m ...

is at least 30 years for males and 25 years for females.

Human interactions

Attacks on humans

Copper sharks attack humans infrequently, but the species places tenth in the number of unprovoked attacks on people. During the tracking period through 2013, the

University of Florida The University of Florida (Florida or UF) is a public university, public land-grant university, land-grant research university in Gainesville, Florida, United States. It is a senior member of the State University System of Florida and a preem ...

attributed 20 attacks to the species. (In comparison,

great white sharks The great white shark (''Carcharodon carcharias''), also known as the white shark, white pointer, or simply great white, is a species of large mackerel shark which can be found in the coastal surface waters of all the major oceans. It is the ...

topped the list, with 279 attacks.) Though large and powerful, the copper shark is not particularly aggressive towards humans unless in the presence of food. Copper Sharks have been known to harass and attack spear fishers in an attempt to steal catches. Copper sharks have bitten several swimmers in

and

, where the species is common. (The species is commonly called bronze whalers in this part of the world.) Fatal attacks attributed to the copper shark (bronze whaler) include the 2014 death of a swimmer in

Tathra, New South Wales Tathra () is a seaside town on the Sapphire Coast found on the South Coast, New South Wales, Australia. As of the , Tathra had a population of 1,675. Nearby points of interest are the Old Tathra Wharf, Mimosa Rocks National Park and Bournda ...

, and the 1976 death of a swimmer in

Te Kaha Te Kaha is a small New Zealand community situated in the Bay of Plenty near Ōpōtiki. The New Zealand Ministry for Culture and Heritage gives a translation of "the rope" for . The full name of Te Kaha is Te-Kahanui-A-Tikirākau. Te Kaha is a ...

. Three out of ten shark attacks in New Zealand are attributed to Copper sharks. Witnesses also attributed a fatal attack in September 2011 in Bunker Bay,

Western Australia Western Australia (WA) is the westernmost state of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Aust ...

to a copper shark. One problem with counting attacks on humans is that the copper shark is so similar to other

s. Victims and witnesses are unlikely to correctly identify which type of genus

Carcharhinus ''Carcharhinus'' is the type genus of the family (biology), family Carcharhinidae, the requiem sharks. One of 12 genera in its family, it contains over half of the species therein. It contains 35 extant and eight extinct species to date, with lik ...

shark is responsible for an attack. Experts trying to confirm shark attacks by species warn that their statistics undercount the number of attacks by requiem sharks like the copper.

Captivity

Like many large, active sharks, this species adapts poorly to captivity; it tends to bump into the sides of its enclosure, and the resulting abrasions then become infected with often fatal consequences.

Fishing

Commercial fisheries Commercial fishing is the activity of catching fish and other seafood for commercial profit, mostly from wild fisheries. It provides a large quantity of food to many countries around the world, but those who practice it as an industry must often p ...

for the copper shark exist off New Zealand, Australia (though the "bronze whaler fishery" of

actually takes mostly dusky sharks), South Africa, Brazil, Uruguay, Argentina, Mexico, and China; it also contributes to the

bycatch Bycatch (or by-catch), in the fishing industry, is a fish or other marine species that is caught unintentionally while fishing for specific species or sizes of wildlife. Bycatch is either the wrong species, the wrong sex, or is undersized or juve ...

of other commercial fisheries across its range. This species is caught in

gillnet Gillnetting is a fishing method that uses gillnets: vertical panels of netting that hang from a line with regularly spaced floaters that hold the line on the surface of the water. The floats are sometimes called "corks" and the line with corks is ...

s and on bottom longlines, and to a much lesser extent in

bottom trawl Bottom trawling is trawling (towing a trawl, which is a fishing net) along the seafloor. It is also referred to as "dragging". The scientific community divides bottom trawling into benthic trawling and Demersal zone, demersal trawling. Benthic tra ...

s and on pelagic longlines. The meat is sold for human consumption. The copper shark is also popular with recreational fishers in Namibia, New Zealand, Australia, South Africa, Argentina, Mexico and California, predominantly by anglers but also by bowfishers and gillnetters. In New Zealand, it is the ''Carcharhinus'' species most frequently caught by sport fishers and sustains a small, summer recreational fishery in northern North Island, that mainly captures pregnant and post-partum females and for the most part practices

tag and release Tag and release is a form of catch and release fishing in which the angler attaches a tag to the fish, records data such as date, time, place, and type of fish on a standardized postcard, and submits this card to a fisheries agency or conser ...

. A tag and release program is also practiced in Namibia.

Conservation

The

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...

(IUCN) has assessed the copper shark as vulnerable worldwide. While the global population is unknown, the long maturation time and low reproductive rate of this species render it highly susceptible to

overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing Fish stocks, fish stock), resu ...

. Regionally, the IUCN has listed this species under

Least Concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...

off Australia, New Zealand, and South Africa, where fisheries are generally well-managed; the local copper shark population for each of those three countries is contained almost entirely within their respective

Exclusive Economic Zone An exclusive economic zone (EEZ), as prescribed by the 1982 United Nations Convention on the Law of the Sea, is an area of the sea in which a sovereign state has exclusive rights regarding the exploration and use of marine natural resource, reso ...

s (EEZ). Reported catches by New Zealand have steadily declined from a peak of 40 tons in 1995/96 to 20 tons 2001/02, though it is uncertain whether this reflects a genuine decline or changing fishing habits. In the eastern Pacific, the copper shark is rare and little fishery information is available, leading to an assessment of

Data Deficient A data deficient (DD) species is one which has been categorized by the International Union for Conservation of Nature (IUCN) as offering insufficient information for a proper assessment of conservation status to be made. This does not necessaril ...

. However, catch declines across all shark and ray species have been documented in the heavily-fished Gulf of California. Off

East Asia East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of Economy of China, China, Economy of Ja ...

, the copper shark has been assessed as Vulnerable; though species-specific data is unavailable, shark populations overall have been decimated in the region. The number of large adults have been too low to sustain targeted fisheries since the 1970s, and most of the current shark catch consists of small juveniles. Additional threats to this species include the

degradation Degradation may refer to: Science * Degradation (geology), lowering of a fluvial surface by erosion * Degradation (telecommunications), of an electronic signal * Biodegradation of organic substances by living organisms * Environmental degradation ...

and

destruction Destruction may refer to: Concepts * Destruktion, a term from the philosophy of Martin Heidegger * Destructive narcissism, a pathological form of narcissism * Self-destructive behaviour, a widely used phrase that ''conceptualises'' certain kin ...

of its coastal nurseries from development,

pollution Pollution is the introduction of contaminants into the natural environment that cause harm. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the component ...

, and

aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. Nelu ...

, mortality from

shark net A shark net is a submerged section of gillnets placed at beaches designed to intercept large marine animals including sharks, with the aim to reduce the likelihood of shark attacks on swimmers. The gillnets form a wall of netting that hangs in t ...

s used to protect beaches in South Africa and Australia, and persecution by fish farmers in southern Australia. The New Zealand

Department of Conservation Department may refer to: * Departmentalization, division of a larger organization into parts with specific responsibility Government and military *Department (administrative division), a geographical and administrative division within a country, ...

has classified the copper shark as "Not Threatened" under the

New Zealand Threat Classification System The New Zealand Threat Classification System is used by the Department of Conservation to assess conservation priorities of species in New Zealand. The system was developed because the IUCN Red List, a similar conservation status system, had s ...

but with the qualifiers "Conservation Dependent", "Data Poor" and "Secure Overseas".

References

External links

''Carcharhinus brachyurus'', Copper shark
a
FishBase''Carcharhinus brachyurus'' (Bronze Whaler, Cocktail Shark, Copper Shark, Narrowtooth Shark, New Zealand Whaler)
a
IUCN Red List
a
Florida Museum of Natural History Ichthyology Department

Species Description of Carcharhinus brachyurus at www.shark-references.com
{{Good article

copper shark The copper shark (''Carcharhinus brachyurus''), bronze whaler, or narrowtooth shark is a species of requiem shark found mostly in temperate latitudes. It is distributed in a number of separate populations in the northeastern and southwestern Atla ...

Fish of the Mediterranean Sea Marine fish of New Zealand Fish of Peru Marine fish of Southern Africa Marine fauna of East Asia Marine fish of Southern Australia Fauna of the Baja California Peninsula Southeastern South American coastal fauna Gulf of Guinea Vulnerable fish

Taxa named by Albert Günther Near threatened animals Near threatened biota of Oceania Near threatened biota of Asia Fish of the Atlantic Ocean Fish of the Indian Ocean Fish of the Pacific Ocean