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''Cananeuretus'' is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22,0 ...
in the Formicidae subfamily
Aneuretinae Aneuretinae is a subfamily of ants consisting of a single extant species, ''Aneuretus simoni'' ( Sri Lankan relict ant), and 9 fossil species. Earlier, the phylogenetic position of ''A. simoni'' was thought to be intermediate between primitive ...
, and is one of two Cretaceous genera of the subfamily. The genus contains a single described species ''Cananeuretus occidentalis'' and is known from one
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
fossil which has been found in North America.


History and classification

''Cananeuretus'' is known from an adult fossil, the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
, specimen number "TMP 8.89.7", and an additional partial worker tentatively assigned to the species. At the time of description, both specimens were residing in the paleontology collections housed at the Royal Tyrell Museum,
Drumheller, Alberta Drumheller is a town on the Red Deer River in the badlands of east-central Alberta, Canada. It is northeast of Calgary and south of Stettler. The Drumheller portion of the Red Deer River valley, often referred to as Dinosaur Valley, has a ...
. The holotype specimen is composed of a mostly complete worker caste adult female which has been preserved as an inclusion in a transparent chunk of clear yellow Canadian amber. The
amber Amber is fossilized tree resin that has been appreciated for its color and natural beauty since Neolithic times. Much valued from antiquity to the present as a gemstone, amber is made into a variety of decorative objects."Amber" (2004). In M ...
specimen was recovered from deposits of the Foremost Formation near Grassy Lake,
Alberta, Canada Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Territo ...
. Canadian amber has been dated to an age of approximately 79–78 million years old, placing it in the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...
of the Late Cretaceous. Analysis of the amber composition indicates it originated as cupressaceous resins which were deposited in lagoons and salt water marshes along the
Western Interior Seaway The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient se ...
. The fossil was first studied by paleoentomologists Michael Engel and David Grimaldi, both of the American Museum of Natural History. Engel and Grimaldi's 2005
type description A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have ...
of the new genus and species was published in the journal ''American Museum Novitates''. The genus name ''Cananeuretus'' was coined as a combination of the genus name "'' Aneuretus''" which is the type genus of the subfamily Aneuretinae, and Canada, in recognition of the country of origin. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''occidentalis'' was based from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
word meaning ''from the west'', a reference the genus being the first Mesozoic aneuretine found in the Western Hemisphere. ''Cananeuretus'' is one of six ant species described from the Grassy Lake site, the others being '' Canapone dentata'', ''
Chronomyrmex medicinehatensis ''Chronomyrmex'' is an extinct genus of ants of the subfamily Dolichoderinae Dolichoderinae is a subfamily of ants, which includes species such as the Argentine ant (''Linepithema humile''), the erratic ant, the odorous house ant, and the ...
'', ''
Eotapinoma macalpini ''Eotapinoma'' is an extinct genus of ants of the subfamily Dolichoderinae Dolichoderinae is a subfamily of ants, which includes species such as the Argentine ant (''Linepithema humile''), the erratic ant, the odorous house ant, and the co ...
'', ''
Haidoterminus cippus ''Haidoterminus'' is an extinct genus of ant in the Formicidae subfamily Haidomyrmecinae, and is one of only nine genera placed in this subfamily. The genus contains a single described species ''Haidoterminus cippus'' and is known from one Lat ...
'', and ''
Sphecomyrma canadensis ''Sphecomyrma'' is an extinct genus of ants which existed in the Cretaceous approximately 79 to 92 million years ago. The first specimens were collected in 1966, found embedded in amber which had been exposed in the cliffs of Cliffwood, New Je ...
''. In the type description, Engel and Grimaldi described ''Cananeuretus'' as a formicomorph genus based the plesiomorphies that are suggested to define the subfamily Aneuretinae. They excluded the genus from
Dolichoderinae Dolichoderinae is a subfamily of ants, which includes species such as the Argentine ant (''Linepithema humile''), the erratic ant, the odorous house ant, and the cone ant. The subfamily presents a great diversity of species throughout the w ...
, which ''Eotapinoma'' belongs to, and separated the two genera based on the well developed sting on ''Cananeuretus'' in combination with an apparent lack of acidopore. ''Cananeuretus'' is the second Mesozoic genus to tentatively be placed into Aneuretinae, after the slightly older '' Burmomyrma'' found in
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. ...
.


Description

''Cananeuretus'' has "primitive" mandibles, each having four distinct teeth present, the basal most being the largest. The compound eyes are small and well developed, being positioned low on the head making observation on the two described specimens hard, and ocelli are absent from both specimens. The twelve segmented antennae are placed in slightly upward positioned antennae sockets and have an elongated scape. The alitrunk is elongated in nature with a narrowing at the front forming a neck like appearance at the front. There are no lobes or projections on the
propodeum The propodeum or propodium is the first abdominal segment in Apocrita Hymenoptera ( wasps, bees and ants). It is fused with the thorax to form the mesosoma. It is a single large sclerite, not subdivided, and bears a pair of spiracles. It is ...
, rather it is smoothly rounded. The legs are slim with a developed grove on the femurs forming trochantellus. The fore-legs have a calcar, or enlarged and modified seta, present. There are small brushes of grouped setae on the interior surfaces of both the protibia and probasitarsus. The gaster is attached to the petiole with an elongated and enlarged connection on the second metasomal segment, while the petiole has a node in the middle. A number of large setae are scattered on the upper surface of the gaster and a large sting is present and extended from the gaster tip.


References

{{Taxonbar, from=Q21078765 Monotypic fossil ant genera Cretaceous insects Fossil taxa described in 2005
A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species). It is one of the modern descendan ...
Aneuretinae Prehistoric insects of North America Canadian amber Taxa named by Michael S. Engel