''Brochocoleus'' is an extinct genus of

beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...

s in the family

Ommatidae The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. Howev ...

, known from the

Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...

to the Early

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...

. 10 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †'' Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia,

Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...

* †'' Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia,

Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...

* †'' Brochocoleus keenani'' Jarzembowski et al., 2013

Weald Clay Weald Clay or the Weald Clay Formation is a Lower Cretaceous sedimentary rock unit underlying areas of South East England, between the North and South Downs, in an area called the Weald Basin. It is the uppermost unit of the Wealden Group of ...

, United Kingdom, Barremian * †'' Brochocoleus maculatus'' (Whalley, 1985)

Charmouth Mudstone Formation The Charmouth Mudstone Formation is a Formation (geology), geological formation in England, dating to the Early Jurassic (Sinemurian–Pliensbachian). It forms part of the lower Lias Group. It is most prominently exposed at its type locality in cl ...

, United Kingdom,

Sinemurian In the geologic timescale, the Sinemurian is an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic epoch (geology), Epoch or series (stratigraphy), Series. It spans the time between 199.5 ±0.3 annu ...

* †'' Brochocoleus maximus'' Jarzembowski et al., 2013

Purbeck Group The Purbeck Group is an Upper Jurassic to Lower Cretaceous lithostratigraphy, lithostratigraphic group (stratigraphy), group (a sequence of rock strata) in south-east England. The name is derived from the district known as the Isle of Purbeck in ...

, United Kingdom,

Berriasian In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...

* †'' Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †'' Brochocoleus rostratus'' Ponomarenko, 1999 Kzyl-Zhar, Kazakhstan,

Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...

* †'' Brochocoleus sacheonensis'' Lee et al., 2023

Jinju Formation The Jinju Formation (), also known as the Dongmyeong Formation () in some literature is an Early Cretaceous geologic Formation (geology), formation in South Korea.Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...

* †'' Brochocoleus sonidensis'' Tang et al., 2024 Damoguaihe Formation,

China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...

* †'' Brochocoleus tobini'' Jarzembowski et al., 2013 Weald Clay, United Kingdom,

Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...

Formerly assigned species

* †''Brochocoleus angustus'' Tan, Ren, & Shih, 2007 Moved to '' Pareuryomma'' * †''Brochocoleus aphaleratus'' (Ponomarenko, 1969) Moved to ''

Allophalerus ''Allophalerus'' is an extinct genus of beetle in the family Ommatidae. It is known from nine species formerly included in the genus ''Tetraphalerus''. Systematics * †''Allophalerus antiquus'' (Ponomarenko 1964) Karabastau Formation, Kazakh ...

'' * †''Brochocoleus applanatus'' Tan & Ren, 2009 Moved to '' Diluticupes'' * †''Brochocoleus caseyi'' Jarzembowski, Wang, & Zheng, 2016 Moved to '' Jarzembowskiops'' * †''Brochocoleus crowsonae'' Jarzembowski, Yan, Wang, & Zhang, 2013 Moved to '' Diluticupes'' * †''Brochocoleus impressus'' (Ren, 1995) Moved to ''Diluticupes'' * †''Brochocoleus indibili'' Soriano & Delclòs, 2006 Considered indeterminate by Kirejtshuk, 2020 * †''Brochocoleus magnus'' Tan & Ren, 2009 Moved to ''Diluticupes'' * †''Brochocoleus minor'' Ponomarenko, 2000 Moved to ''Diluticupes'' * †''Brochocoleus sulcatus'' Tan, Ren, & Shih, 2007 Moved to '' Odontomma'' * †''Brochocoleus validus'' Tan & Ren, 2009 Moved to ''Diluticupes'' * †''Brochocoleus yangshuwanziensis'' Jarzembowski, Yan, Wang, & Zhang, 2013 Moved to ''Diluticupes'' * †''Brochocoleus zhiyuani'' Liu, Tan, Jarzembowski, Wang, Ren, & Pang, 2017 Moved to '' Burmocoleus''

References

Ommatidae Fossil beetle genera {{paleo-beetle-stub