Berothidae

The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings. The family was first named by Anton Handlirsch in 1906. The family consists of 24 genera and 110 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions. Numerous extinct species have also been described. Their ecology is poorly known, but in the species where larval stages have been documented, the larvae are predators of termites.

Systematics

A considerable fossil diversity of beaded lacewings is known from the Late Jurassic onwards, containing numerous genera which are likewise basal or ''incertae sedis.''

;Subfamily Berothimerobiinae
*'' Berothimerobius''
;Subfamily Berothinae
*'' Asadeteva''
*'' Barrowiella''
*'' Berlekrumyia''
*'' Berotha''
* †'' Elektroberotha'' Eocene (Lutetian); Baltic amber, northern Europe
*'' Isoscelipteron''
*'' Lekrugeria''
*'' Lomamyia''
*'' Nodalla''
*'' Podallea''
*'' Quasispermophorella''
*'' Spermophorella''
*'' Spiroberotha''
*'' Stenobiella''
* †'' Xenoberotha'' Green River Formation, United States, Eocene
;Subfamily Cyrenoberothinae
* †'' Aggregataberotha'' (Cenomanian, Burmese amber, Myanmar)
* †'' Araripeberotha'' Cretaceous (Aptian); Crato Formation, Brazil
* †'' Caririberotha'' Cretaceous (Aptian); Crato Formation, Brazil
*'' Cyrenoberotha''
*'' Manselliberotha''
*†'' Microberotha'' (Eocene (Ypresian), Hat Creek amber, Canada)
*'' Ormiscocerus''
*†'' Protoberotha'' (Cenomanian, Burmese amber, Myanmar)
*†'' Sibelliberotha'' (Barremian, Lebanese amber)
*'' Speleoberotha''
;Subfamily † Mesithoninae
* †'' Mesithone'' Callovian/Oxfordian; Karabastau Formation, Kazakhstan; Aptian; Zaza Formation, Turga Formation, Russia
* †'' Oloberotha'' Aptian; Yixian Formation, China
;Subfamily Nosybinae
* †'' Banoberotha'' Cretaceous (Barremian); Lebanese amber, Lebanon
*'' Naizema''
*'' Nosybus''
*'' Tanzanberotha''
;Subfamily Nyrminae
*'' Nyrma''
;Subfamily Protobiellinae
*'' Austroberothella''
*''Protobiella''
;Subfamily Trichomatinae
*'' Trichoberotha''
*'' Trichoma''
;Subfamily ''Incertae sedis''
* †'' Ansoberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Berothone'' Callovian/Oxfordian; Karabastau Formation, Kazakhstan
* †'' Cantabroberotha'' Cretaceous (Albian); Spanish amber, Spain
* †'' Cornoberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Dasyberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Epimesoberotha'' Cretaceous (Berriasian); Lulworth Formation, United Kingdom
* †'' Ethiroberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Haploberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Iceloberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Jersiberotha'' Cenomanian; Burmese amber, Myanmar; Cretaceous (Turonian); New Jersey amber, United States
* †'' Krokhathone'' Jurassic (Callovian/ Oxfordian); Karabastau Formation, Kazakhstan
* †'' Maculaberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Magniberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Nascimberotha'' Turonian; New Jersey amber, United States
* †'' Plesiorobius'' Cretaceous (Campanian); Canadian amber, Canada; Campanian; Taymyr amber, Russia; Cenomanian, unknown formation, Russia
* †'' Proberotha'' Lutetian, Baltic Amber, northern Europe
* †'' Osmyloberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Sinosmylites'' Daohugou, Haifanggou Formation, China, Callovian, Karabastau Formation, Kazakhstan, Callovian/Oxfordian, Ulaan-Ereg Formation, Mongolia, Late Jurassic (Tithonian)
* †'' Systenoberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Telistoberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Xiaoberotha'' Cenomanian; Burmese amber, Myanmar

Formerly included taxa

Subfamily Paraberothinae (now included in Rhachiberothidae)
* †'' Acanthoberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Alboberotha'' Cretaceous (Albian); Charentense amber, France
* †'' Astioberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Chimerhachiberotha'' Barremian; Lebanese amber, Lebanon
* †'' Creagroparaberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Eorhachiberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Kujiberotha'' Cretaceous (Santonian); Kuji amber, Japan
* †'' Micromantispa'' Cenomanian; Burmese amber, Myanmar
* †'' Paraberotha'' Barremian; Lebanese amber, Lebanon
* †'' Pseudosisyra'' Aptian; Zaza Formation, Russia
* †'' Raptorapax'' Barremian; Lebanese amber, Lebanon
* †'' Retinoberotha'' Cenomanian; Bezonnais amber, France,
* †'' Rhachibermissa'' Turonian; New Jersey amber, United States
* †'' Scoloberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Spinoberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Stygioberotha'' Cenomanian; Burmese amber, Myanmar
* †'' Uranoberotha'' Cenomanian; Burmese amber, Myanmar

References

Mantispoidea
Taxa named by Anton Handlirsch
Neuroptera families

{{Neuroptera-stub