Bandfish
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The bandfishes,
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Cepolidae, are 23 species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
es, They are native to the East
Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...
and
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
wherethey dig burrows in sandy or muddy seabeds and eat
zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...
.


Taxonomy

The bandfishes belong to the family Cepolidae, which is the only member of the
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Cepoloidea in the
suborder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...
Percoidei Percoidei is a suborder of bony fishes in the order Perciformes. Many commercially harvested fish species are considered to be contained in this suborder, including the groupers, Serranidae, seabasses and perches. Divisions The following classif ...
of the
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Perciformes Perciformes (), also called the Acanthopteri, is an order or superorder of ray-finned fish in the clade Percomorpha. ''Perciformes'' means " perch-like". Among the well-known members of this group are perches and darters ( Percidae), and als ...
. The family was created in 1810 by the French
naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
Constantine Samuel Rafinesque Constantine Samuel Rafinesque-Schmaltz (; 22 October 178318 September 1840) was a French early 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ult ...
. The placement of the Cepolidae within the Perciformes is not agreed by all authors, some authors place the family with the
Priacanthidae The Priacanthidae, the bigeyes, are a Family (biology), family of 18 species of marine ray-finned fishes. "Catalufa (disambiguation), Catalufa" is an alternate common name for some members of the Priacanthidae. The etymology of the scientific ...
in the order Priacanthiformes, an order which is considered to be ''
incertae sedis or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...
'' within the
series Series may refer to: People with the name * Caroline Series (born 1951), English mathematician, daughter of George Series * George Series (1920–1995), English physicist Arts, entertainment, and media Music * Series, the ordered sets used i ...
Eupercaria Percomorpha () is an extremely large and diverse clade of ray-finned fish. With more than 17,000 known species (including tuna, seahorses, gobies, cichlids, flatfish, wrasse, perches, anglerfish, and pufferfish) known from both marine and fresh ...
.


Subfamilies and genera

The family Cepolidae has 23 species which are arranged into two
subfamilies In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zool ...
and three
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
as follows: * Subfamily Cepolinae Rafinesque, 1815 ** Genus '' Acanthocepola'' Bleeker, 1874 ** Genus ''
Cepola ''Cepola'' is a genus of marine ray-finned fishes belonging to the bandfish family, Cepolidae. The name red bandfish is applied to all members of this genus, but particularly ''C. macrophthalma'', and generally not ''C. australis'', which is a ...
''
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1764
*Subfamily Owstoninae
Jordan Jordan, officially the Hashemite Kingdom of Jordan, is a country in the Southern Levant region of West Asia. Jordan is bordered by Syria to the north, Iraq to the east, Saudi Arabia to the south, and Israel and the occupied Palestinian ter ...
,
Tanaka is the fourth most common Japanese surname. It is typically written with the kanji for . Less common variants include , , , , and . People with the surname *, Japanese musician formerly known as Boku no Lyric no Bōyomi *, Japanese voice actres ...
& Snyder, 1913
** Genus '' Owstonia'' Tanaka, 1908 Some authorities recognise an additional two genera ''Pseudocephala'' and ''Sphenanthias'' but these are regarded as
synonyms A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
of ''Owstonia'' by others.


Etymology

The family name, Cepolidae, is derived from the name coined in 1764 by Linnaeus for the
type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...
, ''Cepola'' and means "little onion", Linnaeus did not explain why he chose this name. It is likely derived from ''cepollam'' or ''cepulam'', which in 1686 was said by
Francis Willughby Francis Willughby (sometimes spelt Willoughby, ) Fellow of the Royal Society, FRS (22 November 1635 – 3 July 1672) was an English ornithology, ornithologist, ichthyology, ichthyologist and mathematician, and an early student of linguistics an ...
to be local names among
Roman Roman or Romans most often refers to: *Rome, the capital city of Italy *Ancient Rome, Roman civilization from 8th century BC to 5th century AD *Roman people, the people of Roman civilization *Epistle to the Romans, shortened to Romans, a letter w ...
fishermen for the similar "''Fierasfer"'', a
pearlfish Pearlfish are marine fish in the ray-finned fish family Carapidae. Pearlfishes inhabit the tropical waters of the Atlantic, Indian, and Pacific Oceans at depths to , along oceanic shelves and slopes. They are slender, elongated fish with no s ...
, to which Linnaeus believed ''
Cepola macrophthalma ''Cepola macrophthalma'' is a species of marine ray-finned fish belonging to the family Cepolidae, the bandfishes. It is found in the eastern Atlantic and Mediterranean from Senegal north to the British Isles. This species is known as the red ba ...
'' was related. As well as this, in 1872
Giovanni Canestrini Giovanni Canestrini (26 December 1835 – 14 February 1900) was an Italian naturalist and biologist and translator who was a native of Revò. Career He initially studied in Gorizia and Meran, then furthered his education in natural sciences at t ...
reported that in
Naples Naples ( ; ; ) is the Regions of Italy, regional capital of Campania and the third-largest city of Italy, after Rome and Milan, with a population of 908,082 within the city's administrative limits as of 2025, while its Metropolitan City of N ...
the common name for ''C. macropthalma'' is ''Pesce cipolia'' meaning "onion fish".


Homonymy

The name Cepolidae Rafinesque, 1815 is a senior
homonym In linguistics, homonyms are words which are either; '' homographs''—words that mean different things, but have the same spelling (regardless of pronunciation), or '' homophones''—words that mean different things, but have the same pronunciat ...
for the family Cepolidae Ihering, 1909 a family of land snails belonging to the superfamily
Helicoidea Helicoidea is a Taxonomy (biology), taxonomic Taxonomic rank, superfamily of air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the order (taxonomy), order Stylommatophora. Taxonomy 2017 taxonomy and l ...
. The case needs be submitted to the
International Commission on Zoological Nomenclature The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries. Orga ...
to resolve the
homonymy In linguistics, homonyms are words which are either; ''homographs''—words that mean different things, but have the same spelling (regardless of pronunciation), or ''homophones''—words that mean different things, but have the same pronunciatio ...
.


Characteristics

The bandfishes are characterised by the possession of a continuous
dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...
with anything between 0 and 4 spines, frequently 3, spines and an
anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...
which has 0 to 2 spines. The
lanceolate The following terms are used to describe leaf plant morphology, morphology in the description and taxonomy (biology), taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade ...
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
is usually joined to the dorsal and anal fins. There is a single pair of postcleithra and there are 6 branchiostegal rays. The
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...
lies along the base of the dorsal fin. The epineural ribs in some of the vertebrae of the
trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure, and the stem of woody plants * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Comput ...
are fused where they are closest to the corresponding
pleural rib The pleural cavity, or pleural space (or sometimes intrapleural space), is the potential space between the pleurae of the pleural sac that surrounds each lung. A small amount of serous pleural fluid is maintained in the pleural cavity to enable ...
s. The overall colour of the body s normally red or pink. The maximum length attained is in '' Cepola rubescens''. They have a large oblique mouth, normally equipped with a single row of thin weakly curved teeth on each jaw, there are no vomerine teeth or palatine teeth. They have large eyes. They have
cycloid scales A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as we ...
which have crenulated edges. The distal pelvic fin rays are unbranched, the remaining proximal four rays are branched.


Distribution

The bandfishes are found in the Eastern Atlantic Ocean, including the
Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...
and the
Indo-West Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the s ...
region, as far south as New Zealand.


Biology

The bandfishes are mostly
fossorial A fossorial animal () is one that is adapted to digging and which lives primarily (but not solely) underground. Examples of fossorial vertebrates are Mole (animal), moles, badgers, naked mole-rats, meerkats, armadillos, wombats, and mole salamand ...
, creating burrows in areas with fine sand or mud substrates. They leave these burrows to feed on
zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...
in the water column. The lay
pelagic The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the sur ...
eggs.


Cultural references

The oldest recipe by a named author involves the preparation of a bandfish. The original recipe book, by Mithaecus, is now lost, but the recipe itself survives thanks to being quoted in the
Deipnosophistae The ''Deipnosophistae'' (, ''Deipnosophistaí'', lit. , where ''sophists'' may be translated more loosely as ) is a work written in Ancient Greek by Athenaeus of Naucratis. It is a long work of Greek literature, literary, Ancient history, h ...
.


References

{{Taxonbar, from=Q806308 Percoidei Taxa named by Constantine Samuel Rafinesque