Bactrian Camel

The Bactrian camel (''Camelus bactrianus''), also known as the Mongolian camel, domestic Bactrian camel or two-humped camel, is a camel native to the steppes of Central Asia. It has two humps on its back, in contrast to the single-humped dromedary. Its population of 2 million exists mainly in the domesticated form. Their name comes from the ancient historical region of Bactria.

Domesticated Bactrian camels have served as pack animals in inner Asia since ancient times. With its tolerance for cold, drought, and high altitudes, it enabled the travel of caravans on the Silk Road. Bactrian camels, whether domesticated or feral, are a separate species from the wild Bactrian camel (''Camelus ferus''), which is the only truly wild (as opposed to feral) species of camelid in the Old World. Domestic Bactrian camels do not descend from wild Bactrian camels, with the two species having split around 1 million years ago.

Taxonomy

The Bactrian camel shares the genus ''Camelus'' with the dromedary (''C. dromedarius'') and the wild Bactrian camel (''C. ferus''). The Bactrian camel belongs to the family Camelidae. The ancient Greek philosopher Aristotle was the first European to describe the camels: in his 4th century BCE ''History of Animals'', he identified the one-humped Arabian camel and the two-humped Bactrian camel. The Bactrian camel was given its current binomial name ''Camelus bactrianus'' by Swedish zoologist Carl Linnaeus in his 1758 publication ''Systema Naturae''.

Though sharing a closer common ancestor with it than with the dromedary, the domestic Bactrian camel does not descend from the wild Bactrian camel, with the two species having diverged hundreds of thousands of years ago, with their mitochondrial genomes estimated to have diverged around 1 million years ago. Genetic evidence suggests that both Bactrian camel species are closely related to the extinct giant camel species '' Camelus knoblochi'' which became extinct around 20,000 years ago, which is equidistant from both living Bactrian camel species.

The Bactrian camel and the dromedary often interbreed to produce fertile offspring. Where the ranges of the two species overlap, such as in northern Punjab, Iran and Afghanistan, the phenotypic differences between them tend to decrease as a result of extensive crossbreeding between them. The fertility of their hybrid has given rise to speculation that the Bactrian camel and the dromedary should be merged into a single species with two varieties. However, a 1994 analysis of the mitochondrial cytochrome b gene revealed that the species display 10.3% divergence in their sequences.

Description

The Bactrian camel is the largest mammal in its native range and is the largest living camel while being shorter at the shoulder than the dromedary. Shoulder height is from with the overall height ranging from , head-and-body length is , and the tail length is . At the top of the humps, the average height is .

Body mass can range from , with males weighing around , and females around . Its long, wooly coat varies in colour from dark brown to sandy beige. A mane and beard of long hair occurs on the neck and throat, with hairs measuring up to long.

The shaggy winter coat is shed extremely rapidly, with huge sections peeling off at once, appearing as if sloppily shorn. The two humps on the back are composed of fat (not water as is sometimes thought). The face is typical of a camelid, being long and somewhat triangular, with a split upper lip. The long eyelashes, along with the sealable nostrils, help to keep out dust in the frequent sandstorms which occur in their natural range. The two broad toes on each foot have undivided soles and are able to spread widely as an adaptation to walking on sand. The feet are very tough, as befits an animal of extreme environments.

Natural habitat

These camels are migratory, and their habitat ranges from rocky mountain massifs to flat steppe, arid desert, (mostly the Gobi Desert), stony plains and sand dunes. Conditions are extremely harsh – vegetation is sparse, water sources are limited and temperatures are extreme. The coat of the Bactrian camel can withstand cold as low as in winter to in summer. The camels' distribution is linked to the availability of water, with large groups congregating near rivers after rain or at the foot of the mountains, where water can be obtained from springs in the summer months, and in the form of snow during the winter.

Life history

Bactrian camels are exceptionally adept at withstanding wide variations in temperature, ranging from freezing cold to blistering heat. They can go without water for months at a time, but when water is available they may drink up to 57 liters at once. When well fed, the humps are plump and erect, but as resources decline, they shrink and lean to the side. When moving faster than a walking speed, they pace, by stepping forwards with both legs on the same side (as opposed to trotting, using alternate diagonals as done by most other quadrupeds). Speeds of up to have been recorded, but they rarely move this fast. Bactrian camels are also said to be good swimmers. Their sense of sight is well developed and their sense of smell is extremely good. The lifespan of Bactrian camels is estimated at up to 50 years, more often 20 to 40 in captivity.

Diet

Bactrian camels are diurnal, sleeping in the open at night and foraging for food during the day. They are primarily herbivorous. With tough mouths that can withstand sharp objects such as thorns, they are able to eat plants that are dry, prickly, salty or bitter, and can ingest virtually any kind of vegetation. When other nutrient sources are not available, these camels may feed on carcasses, gnawing on bones, skin, or various different kinds of flesh. In more extreme conditions, they may eat any material they find, which has included rope, sandals, and even tents. Their ability to feed on a wide range of foods allows them to live in areas with sparse vegetation. The first time food is swallowed, it is not fully chewed. The partly masticated food (called cud) goes into the stomach and later is brought back up for further chewing.

Bactrian camels belong to a fairly small group of animals that regularly eat snow to provide their water needs. Animals living above the snowline may have to do this, as snow and ice can be the only forms of water during winter, and by doing so, their range is greatly enlarged. The latent heat of snow and ice is large compared with the heat capacity of water, forcing animals to eat only small amounts at a time.

Reproduction

Bactrian camels are induced ovulators – they ovulate after insemination (insertion of semen into the vagina); the seminal plasma, not the spermatozoa, induces ovulation. Ovulation occurs in 87% of females after insemination: 66% ovulate within 36 hours and the rest by 48 hours (the same as natural mating). The least amount of semen required to elicit ovulation is about 1.0 ml.

Males during mating time are often quite violent and may bite, spit, or attempt to sit on other male camels. The age of sexual maturity varies, but is usually reached at 3–5 years. Gestation lasts around 13 months. One or occasionally two calves are produced, and the female can give birth to a new calf every other year. Young Bactrian camels are precocial, being able to stand and run shortly after birth, and are fairly large at an average birth weight of . They are nursed for about 1.5 years. The young calf stays with its mother for three to five years, until it reaches sexual maturity, and often helps raise subsequent generations for those years. Wild camels sometimes breed with domesticated or feral camels.
Bactrian camels have been domesticated for thousands of years and are widely bred by pastoralist and agricultural communities across Central Asia, the Middle East, and parts of China and Mongolia. Breeding practices vary by region, but most focus on traits such as hardiness, milk production, load-bearing ability, and temperament. In Mongolia, herders traditionally manage breeding by selectively mating camels during the rutting season, often using older, experienced bulls known for their vigor and fertility. These males are isolated with females to encourage mating and reduce aggression toward other camels.

In Iran and parts of Afghanistan, camel breeding is commonly integrated into mixed farming systems, where camels provide transport and milk alongside other livestock. Semen collection and artificial insemination techniques are increasingly used in breeding programs in Central Asia, particularly in Kazakhstan and Uzbekistan, to improve genetic stock and ensure ovulation timing aligns with optimal conditions for conception.

In China’s Xinjiang region, large-scale camel farms are maintained by both government-supported operations and private breeders, focusing on conserving the genetics of the domesticated Bactrian camel and improving reproductive efficiency. Advances in veterinary care, nutritional supplementation, and controlled breeding environments have increased calving rates in recent decades.

Across these regions, breeding cycles are timed to coincide with favorable seasonal conditions to maximize calf survival. Female camels are usually bred every two years to allow for full recovery after gestation and lactation, although in some cases, higher-frequency breeding is practiced under intensive management systems. Offspring are commonly used in transportation, wool production, or sold in local markets, forming a critical part of the livelihood for many camel-rearing communities.

Climate adaptations

The Bactrian camel has shaggy fur that keeps them warm. They live in areas with extreme temperatures, ranging from intense cold to severe heat. When it gets warm out, their shaggy coat will shed, helping the camel control its body temperature. The Bactrian camel has two humps to store the energy they get from fat. If energy sources become scarce, camels can rely on the energy stored in their humps. Their humps also allow them to hold up to 35 gallons of water at a time.

Evolutionary history

As of the 1980s, a complete range of fossils suggests the first camelids appeared in North America about 30 million years ago, had relatively small body masses, and were adapted to warm climates. By the early Pleistocene (about 2 million years ago), they had already evolved into a form similar to the current Bactrian camel, and many individuals permanently migrated to the opposite end of the Bering Strait in an abrupt fashion, probably as a response to the advancing ice age. The remaining related types of American camelids are now only in South America

Genome

The provides a ''C. bactrianus ferus'' genome using next generation sequencing.
This review...

...cites this study:

Several effective population size studies have been carried out. They show several bottlenecks in both wild and domesticated Bactrians over the past 350,000 years.

Relationship to humans

The Bactrian camel was domesticated circa ~2,500 BCE. The dromedary is believed to have been domesticated between 4000 BCE and 2000 BCE in Arabia. As pack animals, these ungulates are virtually unsurpassed, able to carry at a rate of per day, or over a period of four days.Rahimdad Khan Molai Shedai; Janat ul Sindh 3rd edition 1993; Sindhi Adbi Board Jamshoro, page 20 The species was a mainstay of transportation on the Silk Road. The Bactrian camel was an important aspect in human transportation, they were able to withstand harsh conditions, which made them ideal for long distance travel.  Furthermore, Bactrian camels are frequently ridden, especially in desertified areas. In ancient Sindh, for example, Bactrian camels of two humps were initially used by the rich for riding. The camel was later brought to other areas such as Balochistan and Iran for the same purpose.

Bactrian camels have been the focus of artwork throughout history. For example, westerners from the Tarim Basin and elsewhere were depicted in numerous ceramic figurines of the Chinese Tang dynasty (618–907).

United States imports

Bactrian camels were imported to the U.S. several times in the mid- to late 19th century, both by the U.S. military and by merchants and miners, looking for pack animals sturdier and hardier than horses and mules. Although the camels met these needs, the United States Camel Corps was never considered much of a success. Having brought two shipments of fewer than 100 camels to the U.S., plans were made to import another 1,000, but the US Civil War interrupted this. Most surviving camels of these endeavors, both military and private, were turned loose to survive in the wild. As a result, small feral herds of Bactrian camels existed during the late 19th century in the southwest deserts of the United States.

One United States Camel Corps camel was later sold to Ringling Brothers and was also in several early Hollywood films.

Documentaries

* '' The Story of the Weeping Camel'' is a 2003 Mongolian docudrama about a family of nomadic shepherds trying to get a white calf accepted by his mother, which rejected him after a difficult birth.

Military use

The Indian Army uses these camels to patrol in Ladakh. It was concluded that after carrying out trials and doing a comparative study with a single-humped camel brought from Rajasthan that the double-humped camel is better suited for the task at hand. Colonel Manoj Batra, a veterinary officer of the Indian Army, stated that the double-humped camel "are best suited for these conditions. They can carry loads of at more than which is much more than the ponies that are being used as of now. They can survive without water for at least 72 hours."

Gallery

File:Riding Bactrian camel Nubra.jpg, Riding in Nubra Valley, India
File:Temee.jpg, Mongolia
File:Camels in Kosh-Agachsky District.jpg, alt=Chuya Steppe, Altai Mountains, Russia, Chuya Steppe
File:Bactrian camel in Kyzyllkum.jpg, Bactrian Camels in the Kyzyl Kum Desert
File:Camel Farm in Mongolia 02.jpg, Farm in Central Mongolia
File:SpeakingCamel altColour US7FerrisburghVT 20151011 (39646557970).jpg, Farm in Vermont, US
File:Bactrian camel (Camelus bactrianus) in Tallinn Zoo (March 2023).jpg, alt=In the snow at Tallinn Zoo, Estonia, Tallinn Zoo

Footnotes

See also

*Bactria
*Camel
* List of animals with humps

References

External links

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* – BBC announces discovery of camels in the Gashun Gobi region

{{Authority control

Camels
Mammals of China
Desert fauna
Mammals of India
Mammals of Central Asia
Mammals of Pakistan
Mammals of Mongolia
Livestock
EDGE species
Pack animals
Bactria
Mammals described in 1758
Taxa named by Carl Linnaeus