Australobatrachia ("southern frogs") is a

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

frogs A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough skin texture due to ...

in the suborder

Neobatrachia The Neobatrachia (Neo-Latin ''neo-'' ("new") + ''batrachia'' ("frogs")) are a suborder of the Frog, Anura, the order (biology), order of frogs and toads. This suborder is the most advanced and apomorphic of the three anuran suborders alive toda ...

. It comprises three

families Family (from ) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictability, structure, and safety as ...

of frogs with a

Gondwanan distribution Gondwana ( ; ) was a large landmass, sometimes referred to as a supercontinent. The remnants of Gondwana make up around two-thirds of today's continental area, including South America, Africa, Antarctica, Australia, Zealandia, Arabia, and the ...

, being known from

Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...

Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...

, and

New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...

. Together, they form the

sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to the superfamily

Hyloidea Hyloidea is a superfamily of frogs.R.Alexander Pyron, John J.Wiens, 2011, ''A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians'' Hyloidea accounts for 54% ...

Taxonomy

The common ancestor of all three families inhabited

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...

during the

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...

(about 125 million years ago). By about 100 million years ago, the ancestors of the Calyptocephalellidae diverged from the Myobatrachoidea, as the ancestral Myobatrachoidea moved south, colonizing the Australian continent via the

Antarctic land bridge The Antarctic land bridge was a land bridge connecting the continents of South America, Antarctica, and Australia that existed from the Late Cretaceous to the Late Eocene. The land bridge consisted of the entire continent of Antarctica (at the time ...

. The two families within Myobatrachoidea diverged from each other later in the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...

or during the earliest

Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...

. Australobatrachia contains the following subgroups: *

Calyptocephalellidae The Calyptocephalellidae are a family of toads (although not true members of the Bufonidae) found in Chile containing two living genera, '' Calyptocephalella'' and '' Telmatobufo.'' The genus ''Calyptocephalella'' contains one living species, ...

- Chilean toads (5 species) *

Myobatrachoidea Myobatrachoidea is a superfamily of frogs. It contains two families, both of which are found in Australia, New Guinea, and the Aru Islands. Some sources group these two families into a single family Myobatrachidae. Their closest relatives are ...

Limnodynastidae Limnodynastidae, commonly known as the Australian ground frogs, is a family (biology), family of frogs found in Australia, New Guinea, and the Aru Islands Regency, Aru Islands. They were formerly considered a subfamily of the Myobatrachidae, the ...

- Australian ground frogs (44 species) **

Myobatrachidae Myobatrachidae, commonly known as Australian ground frogs or Australian water frogs, is a family of frogs found in Australia and New Guinea. Members of this family vary greatly in size, from species less than long, to the second-largest frog in ...

- Australian froglets and Australian toadlets (91 species)

References

Neobatrachia Tetrapod unranked clades Taxa named by Darrel Frost {{Neobatrachia-stub