Asterids are a large

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

(

monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...

group) of

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...

s, composed of 17

orders Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * H ...

and more than 80,000 species, about a third of the total flowering plant species. The asterids are divided into the unranked clades

lamiids Asterids are a large clade (monophyletic group) of flowering plants, composed of 17 orders and more than 80,000 species, about a third of the total flowering plant species. The asterids are divided into the unranked clades lamiids (8 orders) and ...

(8 orders) and

campanulids Asterids are a large clade (monophyly, monophyletic group) of flowering plants, composed of 17 Order_(biology), orders and more than 80,000 species, about a third of the total flowering plant species. The asterids are divided into the unranked cl ...

(7 orders), and the single orders Cornales and

Ericales The Ericales are a large and diverse order of flowering plants in the asterid group of the eudicots. Well-known and economically important members of this order include tea and ornamental camellias, persimmon, ebony, blueberry, cranberry, l ...

. Well-known asterids include dogwoods and

hydrangea ''Hydrangea'' ( or ) is a genus of more than 70 species of Flowering plant, flowering plants native plant, native to Asia and the Americas. Hydrangea is also used as the common name for the genus; some (particularly ''Hydrangea macrophylla, H. m ...

s (order Cornales),

tea Tea is an aromatic beverage prepared by pouring hot or boiling water over cured or fresh leaves of '' Camellia sinensis'', an evergreen shrub native to East Asia which probably originated in the borderlands of south-western China and nor ...

blueberries Blueberries are a widely distributed and widespread group of perennial flowering plants with blue or purple berries. They are classified in the section ''Cyanococcus'' with the genus ''Vaccinium''. Commercial blueberries—both wild (lowbush) ...

, cranberries,

kiwifruit Kiwifruit (often shortened to kiwi), or Chinese gooseberry, is the edible berry (botany), berry of several species of woody vines in the genus ''Actinidia''. The most common cultivar group of kiwifruit (Actinidia chinensis var. deliciosa, ...

Brazil nut The Brazil nut (''Bertholletia excelsa'') is a South American tree in the family Lecythidaceae, and it is also the name of the tree's commercially harvested edible seeds. It is one of the largest and longest-lived trees in the Amazon rainforest. ...

s, argan, sapote, and

azalea Azaleas ( ) are flowering shrubs in the genus ''Rhododendron'', particularly the former sections ''Rhododendron sect. Tsutsusi, Tsutsusi'' (evergreen) and ''Pentanthera'' (deciduous). Azaleas bloom in the spring (April and May in the temperate ...

s (order Ericales), sunflowers,

lettuce Lettuce (''Lactuca sativa'') is an annual plant of the family Asteraceae mostly grown as a leaf vegetable. The leaves are most often used raw in Green salad, green salads, although lettuce is also seen in other kinds of food, such as sandwiche ...

, common daisy, yacon,

carrot The carrot ('' Daucus carota'' subsp. ''sativus'') is a root vegetable, typically orange in colour, though heirloom variants including purple, black, red, white, and yellow cultivars exist, all of which are domesticated forms of the wild ...

celery Celery (''Apium graveolens'' Dulce Group or ''Apium graveolens'' var. ''dulce'') is a cultivated plant belonging to the species ''Apium graveolens'' in the family Apiaceae that has been used as a vegetable since ancient times. The original wild ...

parsley Parsley, or garden parsley (''Petroselinum crispum''), is a species of flowering plant in the family Apiaceae that is native to Greece, Morocco and the former Yugoslavia. It has been introduced and naturalisation (biology), naturalized in Eur ...

parsnip The parsnip (''Pastinaca sativa'') is a root vegetable closely related to carrot and parsley, all belonging to the flowering plant family Apiaceae. It is a biennial plant usually grown as an annual. Its long taproot has cream-colored skin an ...

ginseng Ginseng () is the root of plants in the genus ''Panax'', such as South China ginseng (''Panax notoginseng, P. notoginseng''), Korean ginseng (''Panax ginseng, P. ginseng''), and American ginseng (''American ginseng, P. quinquefol ...

, ivies,

holly ''Ilex'' () or holly is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. ''Ilex'' has the most species of any woody dioecious angiosperm genus. The species are evergreen o ...

honeysuckle Honeysuckles are arching shrubs or Vine#Twining vines, twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae. The genus includes 158 species native to northern latitudes in North America, Eurasia, and North Africa. Widely kno ...

, elder, and valerian (clade campanulids),

borage Borage ( or ; ''Borago officinalis''), also known as starflower, is an annual herb in the flowering plant family Boraginaceae native to the Mediterranean region. Although the plant contains small amounts of pyrrolizidine alkaloids, some parts ar ...

, forget-me-nots,

comfrey ''Symphytum'' is a genus of flowering plants in the borage family, Boraginaceae, known by the common name comfrey (pronounced , from the Latin confervere to 'heal' or literally to 'boil together', referring to uses in ancient traditional medicin ...

coffee Coffee is a beverage brewed from roasted, ground coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulating effect on humans, primarily due to its caffeine content, but decaffeinated coffee is also commercially a ...

, frangipani,

gentian ''Gentiana'' () is a genus of flowering plants belonging to the gentian family ( Gentianaceae), the tribe Gentianeae, and the monophyletic subtribe Gentianinae. With over 300 species, it is considered a large genus. Gentians are notable for thei ...

, pong-pong,

oleander ''Nerium oleander'' ( ), commonly known as oleander or rosebay, is a shrub or small tree cultivated worldwide in temperate and subtropical areas as an ornamental and landscaping plant. It is the only species currently classified in the genus ...

, periwinkle,

basil Basil (, ; , ; ''Ocimum basilicum'' (, )), also called great basil, is a culinary herb of the family Lamiaceae (mints). It is a hardiness (plants), tender plant, and is used in cuisines worldwide. In Western cuisine, the generic term "basil" r ...

mint Mint or The Mint may refer to: Plants * Lamiaceae, the mint family ** ''Mentha'', the genus of plants commonly known as "mint" Coins and collectibles * Mint (facility), a facility for manufacturing coins * Mint condition, a state of like-new ...

rosemary ''Salvia rosmarinus'' (), commonly known as rosemary, is a shrub with fragrant, evergreen, needle-like leaves and white, pink, purple, or blue flowers. It is a member of the sage family, Lamiaceae. The species is native to the Mediterranean r ...

, sage,

oregano Oregano (, ; ''Origanum vulgare'') is a species of flowering plant in the mint family, Lamiaceae. It was native to the Mediterranean region, but widely naturalised elsewhere in the temperate climate, temperate Northern Hemisphere. Oregano is a ...

thyme Thyme () is a culinary herb consisting of the dried aerial parts of some members of the genus ''Thymus (plant), Thymus'' of flowering plants in the mint family Lamiaceae. Thymes are native to Eurasia and north Africa. Thymes have culinary, medici ...

lavender ''Lavandula'' (common name lavender) is a genus of 47 known species of perennial flowering plants in the sage family, Lamiaceae. It is native plant, native to the Old World, primarily found across the drier, warmer regions of the Mediterranean ...

, wild dagga,

olive The olive, botanical name ''Olea europaea'' ("European olive"), is a species of Subtropics, subtropical evergreen tree in the Family (biology), family Oleaceae. Originating in Anatolia, Asia Minor, it is abundant throughout the Mediterranean ...

s, ash,

teak Teak (''Tectona grandis'') is a tropical hardwood tree species in the family Lamiaceae. It is a large, deciduous tree that occurs in mixed hardwood forests. ''Tectona grandis'' has small, fragrant white flowers arranged in dense clusters (panic ...

, foxgloves,

lilac ''Syringa'' is a genus of 12 currently recognized species of flowering woody plants in the olive family or Oleaceae called lilacs. These lilacs are native to woodland and scrub from southeastern Europe to eastern Asia, and widely and commonly ...

jasmine Jasmine (botanical name: ''Jasminum'', pronounced ) is a genus of shrubs and vines in the olive family of Oleaceae. It contains around 200 species native to tropical and warm temperate regions of Eurasia, Africa, and Oceania. Jasmines are wid ...

, snapdragons, African violets, butterfly bushes,

sesame Sesame (; ''Sesamum indicum'') is a plant in the genus '' Sesamum'', also called benne. Numerous wild relatives occur in Africa and a smaller number in India. It is widely naturalized in tropical regions around the world and is cultivated for ...

psyllium Psyllium (), or ispaghula (), is the common name used for several members of the plant genus ''Plantago'' whose seeds are used commercially for the production of mucilage. Psyllium is mainly used as a dietary fiber to relieve symptoms of both ...

potato The potato () is a starchy tuberous vegetable native to the Americas that is consumed as a staple food in many parts of the world. Potatoes are underground stem tubers of the plant ''Solanum tuberosum'', a perennial in the nightshade famil ...

es,

eggplant Eggplant (American English, US, Canadian English, CA, Australian English, AU, Philippine English, PH), aubergine (British English, UK, Hiberno English, IE, New Zealand English, NZ), brinjal (Indian English, IN, Singapore English, SG, Malays ...

tomato The tomato (, ), ''Solanum lycopersicum'', is a plant whose fruit is an edible Berry (botany), berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco, potato, and chili peppers. It originate ...

es, chilli peppers,

tobacco Tobacco is the common name of several plants in the genus '' Nicotiana'' of the family Solanaceae, and the general term for any product prepared from the cured leaves of these plants. More than 70 species of tobacco are known, but the ...

petunia ''Petunia'' is a genus of 20 species of flowering plants of South American origin. The popular flower of the same name derived its epithet from the French, which took the word , 'tobacco', from a Tupi–Guarani language. A tender perennial plan ...

morning glory Morning glory (also written as morning-glory) is the common name for over 1,000 species of flowering plants in the family Convolvulaceae, whose taxonomy and systematics remain in flux. These species are distributed across numerous genus, gene ...

, and

sweet potato The sweet potato or sweetpotato (''Ipomoea batatas'') is a dicotyledonous plant in the morning glory family, Convolvulaceae. Its sizeable, starchy, sweet-tasting tuberous roots are used as a root vegetable, which is a staple food in parts of ...

(clade lamiids). Most of the taxa belonging to this clade had been referred to as Asteridae in the

Cronquist system The Cronquist system is a list of systems of plant taxonomy, taxonomic classification system of angiosperms, flowering plants. It was developed by Arthur Cronquist in a series of monographs and texts, including ''The Evolution and Classification of ...

(1981) and as

Sympetalae Sympetally (fused petals) is a flower characteristic that historically was used to classify a grouping of plants termed Sympetalae, but this term has been abandoned in newer molecular based classifications, although the grouping has similarity to ...

in earlier systems. The name asterids (not necessarily capitalised) resembles the earlier

botanical name A botanical name is a formal scientific name conforming to the ''International Code of Nomenclature for algae, fungi, and plants'' (ICN) and, if it concerns a plant cultigen, the additional cultivar or cultivar group, Group epithets must conform t ...

but is intended to be the name of a clade rather than a formal ranked name, in the sense of the ''

ICBN The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN or ICNafp) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all tho ...

''.

History

Genetic analysis carried out after APG II maintains that the sister to all other asterids are the Cornales. A second order that split from the base of the asterids are the

. The remaining orders cluster into two clades, the lamiids and the campanulids. The structure of both of these clades has changed in

APG III The APG III system of flowering plant classification is the third version of a modern, mostly molecular-based, system of plant taxonomy being developed by the Angiosperm Phylogeny Group (APG). Published in 2009, it was superseded in 2016 by a fur ...

. In the APG III system, the following clades were renamed: : euasterids I → lamiids : euasterids II → campanulids

Phylogeny

The phylogenetic tree presented hereinafter has been proposed by the APG IV project.

Subdivision

Lamiids

The lamiid subclade consists of about 40,000 species and account for about 15% of angiosperm diversity, characterized in general by superior

ovaries The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are endocr ...

and corollas with any fusion of the petals () occurring late in the process of development. The major part of lamiid diversity occurs in the group of five orders from Boraginales to Solanales, referred to informally as "core lamiids" (sometimes called Laminae). The remainder of the lamiids are referred to as "basal lamiids", in which Garryales is the

sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to the core lamiids. It has been suggested that the core lamiids radiated from an ancestral line of tropical trees in which the flowers were inconspicuous and the fruit large,

drupaceous In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...

and often single-seeded.

References

Bibliography

External links

*
Asterids
in Stevens, P. F. (2001 onwards)
Angiosperm Phylogeny Website
Version 7, May 2006. {{Taxonbar, from=Q747502 Superasterids Plant unranked clades