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''Artiocetus'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus of early whales belonging to the family Protocetidae. It was a close relative to '' Rodhocetus'' and its tarsals indicate it resembled an artiodactyl.


Etymology

''Artiocetus''' name arises from a combination of cetus and artiodactyl, as this fossil was the first to show that early whales possessed artiodactyl-like ankles. ''Artiocetus'' belongs to the infraorder Cetacea, which includes whales, dolphins and
porpoises Porpoises are a group of fully aquatic marine mammals, all of which are classified under the family Phocoenidae, parvorder Odontoceti (toothed whales). Although similar in appearance to dolphins, they are more closely related to narwhals an ...
. Cetus is a Latinized Greek word literally meaning "sea monster" and is used in biological names to mean "whale". It comes from Ancient Greek κῆτος (kētos), in reference to the sea monster goddess
Ceto Ceto (; grc, Κητώ, Kētṓ, sea monster) is a primordial sea goddess in Greek mythology, the daughter of Pontus and his mother, Gaia. As a mythological figure, she is considered to be one of the most ancient deities, and bore a host of ...
, daughter of
Gaia In Greek mythology, Gaia (; from Ancient Greek , a poetical form of , 'land' or 'earth'),, , . also spelled Gaea , is the personification of the Earth and one of the Greek primordial deities. Gaia is the ancestral mother—sometimes parthenog ...
and Pontus, and said to resemble a gigantic whale or fish. Artiodactyla refers to the mammal order of even-toed ungulates the group containing cattle, deer,
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
s,
giraffe The giraffe is a large African hoofed mammal belonging to the genus ''Giraffa''. It is the tallest living terrestrial animal and the largest ruminant on Earth. Traditionally, giraffes were thought to be one species, ''Giraffa camelopardalis ...
s,
antelope The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia. Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mammals ...
, goats, sheep, pigs and hippopotamuses. If the animal has even number of toes, the weight is borne equally by the third and fourth toe. The shape of the astragalus is another key feature which has a double-pulley structure in artiodactyls, giving the foot greater flexibility.


Description

''Artiocetus clavis'' was a small whale measuring long. It existed in the early Lutetian age (47 million years ago) and is one of the oldest known protocetid archaeocetes. Though the whale may have been primarily aquatic, the discovery of ankle bones lends to the idea that this fossil may have been a transition between sea-based and land-based mammals. While whales eventually returned to the sea, the
anthracotheres Anthracotheriidae is a paraphyletic family of extinct, hippopotamus-like artiodactyl ungulates related to hippopotamuses and whales. The oldest genus, ''Elomeryx'', first appeared during the middle Eocene in Asia. They thrived in Africa and Euras ...
, ancestors of the hippopotamus, are thought to have descended from an ancestor shared with the whale.Supplementary Material: Distinctive characteristics of artiodactyl ankle bones
/ref> Like ''Rodhocetus'', ''Artiocetus'' had limbs comparable to '' Ambulocetus'' but larger fore and hind feet, which were probably webbed. They could probably move on land, but rather clumsily like a modern seal. Protocetidae were the first group of whales to develop tail flukes, which suggests they were quick, agile predators. Though Protocetidae as a family possessed tail flukes, it has been suggested that ''Artiocetus'' did not. Thewissen et al. states that "''Artiocetus'' had a long tail and thus probably lacked a tail fluke".


Fossil finds

Fossils located in 2001 in the Balochistan Province of Pakistan showed that ''Artiocetus'' had both an astragalus and cuboid bone in the ankle (a diagnostic traits of artiodactyls), suggesting that early whales had fore and hind limbs. The distribution of fossils in Indo-Pakistan, Africa, Europe, and North America suggests that this species preferred a warmer sea climate, preferably in the tropics. There is no commonly agreed ancestry of the whale, but they are thought to have evolved from an early group of carnivorous even-toed ungulates. DNA studies have suggested that the hippopotamus is the closest land relative to the whale. ''Artiocetus'' fossils represent intermediate forms between land-living ungulates and whales, lending support to the theory that whales and hippopotami descended from a common ancestor. The discovery of this fossil is important as it helped solidify the theory that whales shared a common ancestor with Artiodactyla. In 2005, an international team of scientists suggested that whales and hippopotami share a common water and terrestrial dwelling ancestor, which lived 50 to 60 million years ago. Two groups emerged from this common ancestor: early cetaceans, which in time returned to the sea permanently, and a large group of superficially pig-like land-based mammals called
anthracotheres Anthracotheriidae is a paraphyletic family of extinct, hippopotamus-like artiodactyl ungulates related to hippopotamuses and whales. The oldest genus, ''Elomeryx'', first appeared during the middle Eocene in Asia. They thrived in Africa and Euras ...
. The only surviving descendants of anthracotheres are the
common Common may refer to: Places * Common, a townland in County Tyrone, Northern Ireland * Boston Common, a central public park in Boston, Massachusetts * Cambridge Common, common land area in Cambridge, Massachusetts * Clapham Common, originally com ...
and pygmy hippopotamuses.


References


External links

* {{Taxonbar, from=Q4801186 Protocetidae Fossil taxa described in 2001 Prehistoric cetacean genera Eocene mammals of Asia