Ark Clam
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Ark clam is the
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...
for a family of small to large-sized saltwater clams or marine
bivalve Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class (biology), class of aquatic animal, aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed b ...
mollusc Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
s in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Arcidae. Generally less than 80 mm long, ark clams vary both in shape and size. They number about 200 species worldwide. The group is known as "ark shells" because species such as '' Arca'' have a large flat area between the umbones which, in an undamaged shell, somewhat resembles a deck, with the rest of the shell perhaps illustrating an ancient wooden boat such as Noah's ark is thought to have been. The thick, ridged shells of ark clams are often white, cream or tan, but in some species, the shell is striped with, tinted with, or completely colored, a rich brown. In life the shell of most species has a top shell layer that is thick brown
periostracum The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods an ...
affixed to the harder calcareous part of the shell. In some species such as '' Barbatia'', this outer horny covering is tufted at the end of the shell into something that resembles a beard, hence the name ''Barbatia'' or bearded one. The thick outer skin or
periostracum The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods an ...
of an ark clam can act as
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
, such that the shells can sometimes look like stones when lying on the bottom. All ark shells have a long straight hinge line with a single row of numerous small and unspecialized "teeth". This is known as a "taxodont dentition" and represents an ancient ancestor. This kind of hinge line is also found in the bivalve families Glycymerididae, Nuculidae and Nuculanidae. Ark clams are distinct from other clams in having red blood pigments (
hemoglobin Hemoglobin (haemoglobin, Hb or Hgb) is a protein containing iron that facilitates the transportation of oxygen in red blood cells. Almost all vertebrates contain hemoglobin, with the sole exception of the fish family Channichthyidae. Hemoglobin ...
) that facilitate the transport of
oxygen Oxygen is a chemical element; it has chemical symbol, symbol O and atomic number 8. It is a member of the chalcogen group (periodic table), group in the periodic table, a highly reactivity (chemistry), reactive nonmetal (chemistry), non ...
to their tissues and enable them to inhabit more hypoxic environments. Ark clams reach reproductive maturity when they are about twelve months old and about 20 mm long. The
spawning Spawn is the Egg cell, eggs and Spermatozoa, sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of freely releasing eggs and sperm into a body of water (fresh or marine); the physical act is ...
cycle typically begins in the rainy season. Ark Clams are ''broadcast spawners'', that is, eggs and sperm are released into open water where fertilization occurs. The fertilized eggs develop rapidly into
planktonic Plankton are the diverse collection of organisms that drift in water (or air) but are unable to actively propel themselves against currents (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they pro ...
larvae A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect developmental biology, development such as insects, some arachnids, amphibians, or cnidarians typical ...
that drift with ocean currents for eight to ten days during which 99.9% of the larvae are consumed or perish. Eventually the survivors settle to suitable sites of the sea floor where they develop into juvenile clams. Only one percent of these juveniles will survive to become a mature adult. Ark Clams have a maximum life span of about six years.


Human use

Ark clams are edible mollusks that have been consumed since pre-historic times and are still consumed today. Numerous recipes for ark clams have been published. Recreational foragers should follow seafood safety guidelines and shellfish harvesting restrictions, and should be aware of any harmful algal blooms that may contaminate shellfish with biotoxins. Shellfish collected in urban areas should not be eaten raw. '' Tegillarca granosa'' was used as a food by Indigenous peoples living on the northern Australian coastline through at least the past ~4500 years, with extensive evidence preserved in the form of
shell mound A midden is an old dump for domestic waste. It may consist of animal bones, human excrement, botanical material, mollusc shells, potsherds, lithics (especially debitage), and other artifacts and ecofacts associated with past human occup ...
sites. Large ark clams, such as '' Arca zebra'', are commonly used as bait, as well as food, throughout the
Caribbean The Caribbean ( , ; ; ; ) is a region in the middle of the Americas centered around the Caribbean Sea in the Atlantic Ocean, North Atlantic Ocean, mostly overlapping with the West Indies. Bordered by North America to the north, Central America ...
. In Japan, red Ark clams, called Akagai, are used in
sushi is a traditional Japanese dish made with , typically seasoned with sugar and salt, and combined with a variety of , such as seafood, vegetables, or meat: raw seafood is the most common, although some may be cooked. While sushi comes in n ...
or
sashimi is a Japanese cuisine, Japanese delicacy consisting of fresh raw fish or Raw meat, meat sliced into thin pieces and often eaten with soy sauce. Origin The word ''sashimi'' means 'pierced body', i.e., "wikt:刺身, 刺身" = ''sashimi'', whe ...
. Some ark clams species, such as the blood cockle ('' Anadara granosa'', a.k.a. '' Tegillarca granosa'') are raised in
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. Nelu ...
, e.g. in the estuaries of China's
Fujian Fujian is a provinces of China, province in East China, southeastern China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capital is Fuzhou and its largest prefe ...
coast. In the U.S. limited quantities of wild ark clams have been harvested in
North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...
and
Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...
for ethnic markets and aquaculture has been explored. In the south Pacific region, ''Ark'' clams are still gathered by indigenous people as an important subsistence food. Ark clams are harvested, mostly by women, by
gleaning Gleaning is the act of collecting leftover crops in the field after harvest. During harvest, there is food that is left or missed often because it does not meet store standards for uniformity. Sometimes, fields are left because they were not ec ...
intertidal zones. To maintain the Ark Clam fishery, several communities in
Fiji Fiji, officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists of an archipelago of more than 330 islands—of which about ...
are imposing a minimum size limit of 3 cm, closures during spawning periods, and establishing "no-take" areas.


Genera

Genera within the family Arcidae include: * '' Acar''
Gray Grey (more frequent in British English) or gray (more frequent in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning that it has no chroma. It is the color of a cloud-covered s ...
, 1857
* ''
Anadara ''Anadara'' is a genus of seawater, saltwater bivalves, ark clams, in the family (biology), family Arcidae. It is also called ''Scapharca''. This genus is known in the fossil record from the Cretaceous period to the Quaternary period (age range ...
''
Gray Grey (more frequent in British English) or gray (more frequent in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning that it has no chroma. It is the color of a cloud-covered s ...
, 1847
* '' Arca''
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of ''Systema Naturae'', the starting point of modern zoologic ...
* '' Barbatia''
Gray Grey (more frequent in British English) or gray (more frequent in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning that it has no chroma. It is the color of a cloud-covered s ...
, 1847
* '' Bathyarca'' Kobelt, 1891 * '' Bentharca'' Verrill and Bush, 1898 * '' Larkinia'' Reinhart, 1935 * '' Samacar'' Iredale, 1936 * '' Senilia''
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of ''Systema Naturae'', the starting point of modern zoologic ...
* '' Tegillarca'' Iredale, 1939 * '' Palestinarca''†


References


External links


ITIS


{{Authority control Clams Edible molluscs