Arion Fuscus
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''Arion fuscus'', also known as the "dusky arion", is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of small air-breathing land
slug Slug, or land slug, is a common name for any apparently shell-less Terrestrial mollusc, terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced ...
, a
terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
pulmonate Pulmonata or pulmonates is an informal group (previously an order, and before that, a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group inclu ...
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000  extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Arionidae Arionidae, common name the "roundback slugs" or "round back slugs" are a taxonomic family of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Arionoidea. Distribution The distribution of this family of s ...
, the roundback slugs. ''Arion subfuscus'' (Draparnaud, 1805) is considered by some authors to be a partial
synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
for ''Arion fuscus''. ''Arion subfuscus'' (Draparnaud, 1805) and ''A. fuscus'' (Müller, 1774) have very similar morphology and have overlapping distributional ranges in northwestern Europe. ''Arion fuscus'' is widespread throughout Central, North and East Europe whereas ''A. subfuscus'' is restricted to West Europe. (For practical purposes the two are synonyms in Germany and in the Czech Republic.) The two taxa are referred to as the
species complex In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
''Arion subfuscus''/''fuscus'' both members having several evolutionary lineages. They are only separable by
alloenzyme Alloenzymes (or also called allozymes) are variant forms of an enzyme which differ structurally but not functionally from other allozymes coded for by different alleles at the same locus. These are opposed to isozymes, which are enzymes that p ...
analysis and gonad type.


Description

A 50–70 mm roundback slug. It is brown in colour, ranging from black-brown through olive-brown to orange and bright reddish-orange. Darker lateral bands are present. Dark pigment reaches under the pneumostome from back to front. The foot fringe is pale with thin lines. The sole is uniformly pale/translucent.The tentacles are pale or dark. It does not contract into a bell shape when stimulated. The Dusky Arion differs from other ''Arion'' species in its yellow-orange body mucus. Genitalia:Atrium about half diameter of spermatheca, spermatheca spherical, oviduct large and swollen like in ''
Arion lusitanicus ''Arion lusitanicus'', also known by its common name Portuguese slug, is a species of air-breathing land slug, a terrestrial molluscs, terrestrial pulmonate gastropod mollusc, mollusk in the family Arionidae. Distribution ''Arion lusitanicus'' ...
''


Distribution

This species occurs in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
an and
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
n countries and islands including: *
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
*
Bulgaria Bulgaria, officially the Republic of Bulgaria, is a country in Southeast Europe. It is situated on the eastern portion of the Balkans directly south of the Danube river and west of the Black Sea. Bulgaria is bordered by Greece and Turkey t ...
*
Czech Republic The Czech Republic, also known as Czechia, and historically known as Bohemia, is a landlocked country in Central Europe. The country is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the south ...
- least concern (LC) *
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
*
Netherlands , Terminology of the Low Countries, informally Holland, is a country in Northwestern Europe, with Caribbean Netherlands, overseas territories in the Caribbean. It is the largest of the four constituent countries of the Kingdom of the Nether ...
Arion subfuscus — Anemoon
/ref> *
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...
*
Slovakia Slovakia, officially the Slovak Republic, is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the west, and the Czech Republic to the northwest. Slovakia's m ...
*
Great Britain Great Britain is an island in the North Atlantic Ocean off the north-west coast of continental Europe, consisting of the countries England, Scotland, and Wales. With an area of , it is the largest of the British Isles, the List of European ...
*
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
*
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
Wiktor A., De-niu C. & Wu M. (2000). "Stylommatophoran slugs of China (Gastropoda: Pulmonata) – prodromus". ''Folia Malacologica'' 8(1): 3-35. * and other areas


Habitat

This slug lives under fallen logs and under bark in
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
areas, but it also lives in hedges,
garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate bot ...
s,
pasture Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Types of pasture Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, c ...
s and
dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, flat ...
s.


References


Further reading

* Pinceel J., Joardens K., Van Houtte N., De Winter A.J. & Backeljau T. 2004: Molecular and morphological data reveal cryptic taxonomic diversity in the terrestrial slug complex Arion subfuscus/fuscus (Mollusca, Pulmonata, Arionidae) in continental north-west Europe. Biological Journal of the Linnean Society 83: 23–38. * Pinceel J., Joardens K., Pfenninger M. & Backeljau T. 2005: Rangewide phylogeography of a terrestrial slug in Europe: evidence for Alpine refugia and rapid colonization after the Pleistocene glaciations. Molecular Ecology 14: 1133–1150. * * "Activity and Ecological Distribution of the Slug, ''Arion subfuscus'' (Draparnaud) (Stylommatophora, Arionidae)". * "Relations between forest management and slug assemblages (Gastropoda) of deciduous regrowth forests".


External links


''Arion subfuscus''
at
Animalbase AnimalBase is a project brought to life in 2004 and is maintained by the University of Göttingen, Germany. The goal of the AnimalBase project is to digitize early zoological literature, provide copyright-free open access to zoological works, and p ...
taxonomy,short description, distribution, biology,status (threats), images
''Arion subfuscus''
at
Encyclopedia of Life The Encyclopedia of Life (EOL) is a free, online encyclopedia intended to document all of the 1.9 million living species known to science. It aggregates content to form "pages" for every known species. Content is compiled from existing trusted ...
Arion (gastropod) Gastropods of Europe Gastropods of Asia Fauna of England Gastropods described in 1774 Taxa named by Otto Friedrich Müller {{Arionidae-stub