Anomura (sometimes Anomala) is a group of decapod crustaceans, including hermit crabs and others. Although the names of many anomurans include the word ''crab'', all true

crab Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the ...

s are in the

sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...

to the Anomura, the Brachyura (the two groups together form the

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

Meiura).

Description

The name Anomura derives from an old classification in which reptant decapods were divided into Macrura (long-tailed), Brachyura (short-tailed) and Anomura (differently-tailed). The alternative name Anomala reflects the unusual variety of forms in this group; whereas all crabs share some obvious similarities, the various groups of anomurans are quite dissimilar. The group has been moulded by several instances of carcinisation – the development of a crab-like body form. Thus, the king crabs (Lithodidae), porcelain crabs (Porcellanidae) and hairy stone crab (Lomisidae) are all separate instances of carcinisation. As decapods (meaning ''ten-legged''), anomurans have ten

pereiopod The decapod (crustaceans such as a crab, lobster, shrimp or prawn) is made up of 20 body segments grouped into two main body parts: the cephalothorax and the pleon (abdomen). Each segment may possess one pair of appendages, although in various g ...

s, but the last pair of these is reduced in size, and often hidden inside the gill chamber (under the

carapace A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...

) to be used for cleaning the gills. Since this arrangement is very rare in true crabs (for example, the small family Hexapodidae), a "crab" with only eight visible pereiopods is generally an anomuran.

Evolution

The infraorder Anomura belongs to the group Reptantia, which consists of the walking/crawling decapods ( lobsters and crabs). There is wide acceptance from morphological and

molecular A molecule is a group of two or more atoms held together by attractive forces known as chemical bonds; depending on context, the term may or may not include ions which satisfy this criterion. In quantum physics, organic chemistry, and bioche ...

data that Anomura and Brachyura (true crabs) are sister taxa, together making up the

Meiura. The cladogram below shows Anomura's placement within the larger

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...

Decapoda The Decapoda or decapods (literally "ten-footed") are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crabs, lobsters, crayfish, shrimp and prawns. Most decapods are scavengers. The order is esti ...

, from analysis by Wolfe ''et al.'' (2019). Some of the internal relationships within Anomura can be shown in the cladogram below, which shows Hippidae as sister to Paguroidea, and resolves Parapaguridae outside of Paguroidea:

Classification

The infraorder Anomura contained seven extant superfamilies: The oldest fossil attributed to Anomura is ''

Platykotta ''Platykotta akaina'' is a species of decapod crustacean from the Triassic of the United Arab Emirates. It is the oldest known fossil from the infraorder Anomura, and is most closely related to '' Eocarcinus praecursor''. Stratigraphy The holo ...

'', from the

Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ...

–

Rhaetian The Rhaetian is the latest age of the Triassic Period (in geochronology) or the uppermost stage of the Triassic System (in chronostratigraphy). It was preceded by the Norian and succeeded by the Hettangian (the lowermost stage or earliest age ...

(Late Triassic) Period in the United Arab Emirates.

References

External links

* * {{Taxonbar, from=Q526867 Arthropod infraorders Norian first appearances Extant Late Triassic first appearances