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The angle shades (''Phlogophora meticulosa'') is a
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
of the family
Noctuidae The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family (biology), family of moths. Taxonomically, they are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly ...
. The species was first described by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in his 1758 10th edition of ''Systema Naturae''. It is distributed throughout Europe as far east as the
Urals The Ural Mountains ( ),; , ; , or simply the Urals, are a mountain range in Eurasia that runs north–south mostly through Russia, from the coast of the Arctic Ocean to the river Ural (river), Ural and northwestern Kazakhstan.
and also in the
Azores The Azores ( , , ; , ), officially the Autonomous Region of the Azores (), is one of the two autonomous regions of Portugal (along with Madeira). It is an archipelago composed of nine volcanic islands in the Macaronesia region of the North Atl ...
, in
Algeria Algeria, officially the People's Democratic Republic of Algeria, is a country in the Maghreb region of North Africa. It is bordered to Algeria–Tunisia border, the northeast by Tunisia; to Algeria–Libya border, the east by Libya; to Alger ...
, and in
Asia Minor Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
,
Armenia Armenia, officially the Republic of Armenia, is a landlocked country in the Armenian Highlands of West Asia. It is a part of the Caucasus region and is bordered by Turkey to the west, Georgia (country), Georgia to the north and Azerbaijan to ...
, and
Syria Syria, officially the Syrian Arab Republic, is a country in West Asia located in the Eastern Mediterranean and the Levant. It borders the Mediterranean Sea to the west, Turkey to Syria–Turkey border, the north, Iraq to Iraq–Syria border, t ...
. It is strongly migratory.


Description

This species has a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...
of 45–52 mm and the forewings are very distinctively shaped with a sharply pointed apex. The common name is derived from the characteristic markings on the forewings: the base colour is buffish, brown towards the , and is marked with a bold V-shaped pink-and-green marking. Despite this bright colouring, the angular markings provide excellent disruptive patterning camouflage. The hindwings are whitish with darker venation.


Technical description and variation

This moth has a wingspan of 45–52 mm. Forewing whitish ochreous, the base and costal area extensively pinkish; a triangular space on inner margin before the inner line, the terminal area beyond submarginal line, and a costal shade beyond outer line olive greenish; central area dark green, pinkish towards costa, triangular in shape, the blunt apex resting on inner margin; the three stigmata more or less rosy green, the two upper with pale lateral edges; outer line double and angled outwards on vein 5; submarginal line preceded by a blackish-green lunule between veins 6 and 7; fringe rufous green, blackish along the excision below vein 4; hindwing pale ochreous, with the discal spot, veins, and often the whole inner half tinged with greenish fuscous; dark outer and double submarginal lines, the latter often forming a grey band below vein 4; ab. ''roseobrunnea'' ab. nov arrenfrom
São Jorge Island São Jorge (; Portuguese language, Portuguese for 'Saint George') is an island in the central group of the Azores archipelago and part of the autonomous region of Portugal. Separated from its nearest neighbours (Pico Island, Pico and Faial Isl ...
in the Azores, has the central triangle rich red brown tinged with fulvous, the whole wing reddish tinged, and the green shades all strongly mixed with reddish, the metathorax and dorsal tufts also being deep fulvous instead of green.


Biology

This species flies mainly at night and is attracted to light and sugar. It may often be seen during the day at rest on fences and garden foliage. The larva is dull green or pinkish brown, thickly dotted with pale; dorsal line whitish, interrupted; lateral lines narrowly whitish; segments 4–11 with oblique darker lateral stripes. The
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
is green or brown with reddish spots along the sides and faint dark chevrons along the back. It feeds on a wide variety of plants (see list below). This species overwinters as a larva. There is a similar related moth called the
small angle shades The small angle shades (''Euplexia lucipara'') is a moth of the family Noctuidae. It is distributed throughout the Palearctic. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. As the common name ...
, ''Euplexia lucipara''. File:Phlogophora meticulosa 14-12-2005 18-31-02.JPG, Green and brown larvae File:Phlogophora.meticulosa.6964.jpg, Caterpillar File:Phlogophora.meticulosa.pupa.6970.jpg, Pupa File:Phlogophora meticulosa-02 (xndr).jpg, Close-up of imago File:Phlogophora meticulosa-03 (xndr).jpg File:Phlogophora meticulosa 1-3-2006 14-39-16.JPG, Moth File:Phlogophora meticulosa 20-1-2006 17-10-57.JPG, Caterpillar (brown variant) on '' Aeonium'' File:Phlogophora meticulosa 23-12-2005 13-23-36.JPG, Caterpillar (green variant) on ''Aeonium''


Recorded food plants

*'' Aeonium'' *'' Anemone'' *''
Apium ''Apium'' is a genus, as currently circumscribed by Plants of the World Online, of 12 species of flowering plants in the family Apiaceae, with an unusual highly disjunct distribution with one species in the temperate Northern Hemisphere in the We ...
'' – celery *''
Basil Basil (, ; , ; ''Ocimum basilicum'' (, )), also called great basil, is a culinary herb of the family Lamiaceae (mints). It is a hardiness (plants), tender plant, and is used in cuisines worldwide. In Western cuisine, the generic term "basil" r ...
'' *''Beta'' –
beet The beetroot (British English) or beet (North American English) is the taproot portion of a '' Beta vulgaris'' subsp. ''vulgaris'' plant in the Conditiva Group. The plant is a root vegetable also known as the table beet, garden beet, dinner ...
*''Betula'' –
birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech- oak family Fagaceae. The genus ''Betula'' contains 3 ...
*''
Brassica ''Brassica'' () is a genus of plants in the cabbage and mustard family (Brassicaceae). The members of the genus are informally known as cruciferous vegetables, cabbages, mustard plants, or simply brassicas. Crops from this genus are sometim ...
'' *''
Broccoli Broccoli (''Brassica oleracea'' var. ''italica'') is an edible green plant in the Brassicaceae, cabbage family (family Brassicaceae, genus ''Brassica'') whose large Pseudanthium, flowering head, plant stem, stalk and small associated leafy gre ...
'' *''Centranthus'' – red valerian *''
Chrysanthemum Chrysanthemums ( ), sometimes called mums or chrysanths, are flowering plants in the Asteraceae family. They are native to East Asia and northeastern Europe. Most species originate from East Asia, and the center of diversity is in China. Co ...
'' *''Cynara'' – globe artichoke *''
Dahlia ''Dahlia'' ( , ) is a genus of bushy, tuberous, herbaceous perennial plants native to Mexico and Central America. Dahlias are members of the Asteraceae (synonym name: Compositae) family of dicotyledonous plants, its relatives include the sun ...
'' *''Fragaria'' –
strawberry The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown Hybrid (biology), hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit ...
*''Geranium'' *''Hedera'' – ivy *''
Helianthus ''Helianthus'' () is a genus comprising around 70 species of annual and perennial flowering plants in the daisy family Asteraceae commonly known as sunflowers. Except for three South American species, the species of ''Helianthus'' are native to ...
'' – sunflower *'' Humulus'' – hop *'' Lactuca'' – lettuce *''
Lamium ''Lamium'' (dead-nettles) is a genus of about 30 species of flowering plants in the family Lamiaceae, of which it is the type genus. They are all herbaceous plants native to Eurasia and northern Africa, with several widely naturalised across muc ...
'' – deadnettle *''
Malus ''Malus'' ( or ) is a genus of about 32–57 species of small deciduous trees or shrubs in the family Rosaceae, including the domesticated orchard apple, crab apples (sometimes known in North America as crabapples) and wild apples. The genus i ...
'' – apple *''Nicotiana'' –
tobacco Tobacco is the common name of several plants in the genus '' Nicotiana'' of the family Solanaceae, and the general term for any product prepared from the cured leaves of these plants. More than 70 species of tobacco are known, but the ...
*''
Ocimum basilicum Basil (, ; , ; ''Ocimum basilicum'' (, )), also called great basil, is a culinary herb of the family Lamiaceae (mints). It is a hardiness (plants), tender plant, and is used in cuisines worldwide. In Western cuisine, the generic term "basil" r ...
'' -
basil Basil (, ; , ; ''Ocimum basilicum'' (, )), also called great basil, is a culinary herb of the family Lamiaceae (mints). It is a hardiness (plants), tender plant, and is used in cuisines worldwide. In Western cuisine, the generic term "basil" r ...
*''
Pelargonium ''Pelargonium'' () is a genus of flowering plants that includes about 280 species of perennial plant, perennials, succulent plant, succulents, and shrubs, common name, commonly called geraniums, pelargoniums, or storksbills. ''Geranium'' is also ...
'' *''
Prunus ''Prunus'' is a genus of flowering plant, flowering trees and shrubs from the family (biology), family Rosaceae. The genus includes plums, cherries, peaches, nectarines, apricots and almonds (collectively Drupe, stonefruit). The genus has a cosm ...
'' *''Pyrus'' –
pear Pears are fruits produced and consumed around the world, growing on a tree and harvested in late summer into mid-autumn. The pear tree and shrub are a species of genus ''Pyrus'' , in the Family (biology), family Rosaceae, bearing the Pome, po ...
*''Quercus'' –
oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
*''
Rubus ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. ...
'' – bramble *''
Rumex The docks and sorrels, genus ''Rumex'', are a genus of about 200 species of annual, biennial, and perennial herbs in the buckwheat family, Polygonaceae. Members of this genus are very common perennial herbs with a native almost worldwide distri ...
'' – dock *''
Senecio ''Senecio'' is a genus of flowering plants in the daisy family (Asteraceae) that includes ragworts and groundsels. Variously circumscribed taxonomically, the genus ''Senecio'' is one of the largest genera of flowering plants. Description Mo ...
'' *''
Solanum ''Solanum'' is a large and diverse genus of flowering plants, which include three food crops of high economic importance: the potato, the tomato and the eggplant (aubergine, brinjal). It is the largest genus in the nightshade family Solana ...
'' *''Spinacia'' –
spinach Spinach (''Spinacia oleracea'') is a leafy green flowering plant native to Central Asia, Central and Western Asia. It is of the order Caryophyllales, family Amaranthaceae, subfamily Chenopodioideae. Its leaves are a common vegetable consumed eit ...
*''
Stellaria ''Stellaria'' is a genus of about 190 species of flowering plants in the family Caryophyllaceae, with a cosmopolitan distribution. Common names include starwort, stitchwort and chickweed. Description ''Stellaria'' species are relatively small ...
'' *''
Urtica ''Urtica'' is a genus of flowering plants in the family Urticaceae. Many species have stinging hairs and may be called nettles or stinging nettles (the latter name applying particularly to ''Urtica dioica, U. dioica''). The generic name ''Urtica' ...
'' – nettle *'' Vicia'' *''Vitis'' –
grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus ''Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began approximately 8,0 ...
*''
Mentha ''Mentha'', also known as mint (from Greek , Linear B ''mi-ta''), is a genus of flowering plants in the mint family, Lamiaceae. It is estimated that 13 to 24 species exist, but the exact distinction between species is unclear. Hybridization oc ...
'' -
mint Mint or The Mint may refer to: Plants * Lamiaceae, the mint family ** ''Mentha'', the genus of plants commonly known as "mint" Coins and collectibles * Mint (facility), a facility for manufacturing coins * Mint condition, a state of like-new ...
See Robinson, G. S. et al.


References

* Chinery, Michael ''Collins Guide to the Insects of Britain and Western Europe'', 1986 (Reprinted 1991) * Skinner, Bernard ''Colour Identification Guide to Moths of the British Isles'', 1984


External links

* * Taxonomy
''Lepiforum e.V.''
{{Taxonbar , from=Q768921 Noctuinae Moths described in 1758 Moths of Asia Moths of Europe Moths of Iceland Moths of the Middle East Taxa named by Carl Linnaeus