Angarosphecidae
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Angarosphecidae is an extinct family of
Mesozoic The Mesozoic Era is the Era (geology), era of Earth's Geologic time scale, geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceous Period (geology), Periods. It is characterized by the dominance of archosaurian r ...
and early Cenozoic
wasps A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...
in the superfamily
Apoidea The superfamily Apoidea is a major group (of over 30 000 species) within the Hymenoptera, which includes two traditionally recognized lineages, the "sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from ...
.


Taxonomy

* '' Angarosphex'' (synonyms ''Calobaissodes'' , ''Mataeosphex'' , ''Palaeapis'' , ''Shandongodes'' **''An. alethes'' -
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
, Myanmar **''An. baektoensus'' - Barremian/Aptian, Sinuiju Formation, North Korea **''An. beiboziensis'' -
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
Laiyang Formation, China **''An. bleachi'' -
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
, Clockhouse Brickworks,
Weald Clay Weald Clay or the Weald Clay Formation is a Lower Cretaceous sedimentary rock unit underlying areas of South East England, between the North and South Downs, in an area called the Weald Basin. It is the uppermost unit of the Wealden Group of ...
, United Kingdom **''An. consensus'' -
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
, Clockhouse Brickworks,
Weald Clay Weald Clay or the Weald Clay Formation is a Lower Cretaceous sedimentary rock unit underlying areas of South East England, between the North and South Downs, in an area called the Weald Basin. It is the uppermost unit of the Wealden Group of ...
, United Kingdom **''An. goldringi'' -
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
, Auclaye Brickworks,
Weald Clay Weald Clay or the Weald Clay Formation is a Lower Cretaceous sedimentary rock unit underlying areas of South East England, between the North and South Downs, in an area called the Weald Basin. It is the uppermost unit of the Wealden Group of ...
, United Kingdom **''An. lithodes'' -
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
Laiyang Formation,
Yixian Formation The Yixian Formation (; formerly Romanization of Chinese, transcribed as Yihsien Formation or Yixiang Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the Barremian stage of the Early Cretaceous. I ...
, China **''An. lithographicus'' -
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
/
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
La Pedrera de Rúbies Formation, Spain **''An. magnus'' - Barremian
Crato Formation The Crato Formation is a geologic formation (stratigraphy), formation of Early Cretaceous (Aptian) age in northeastern Brazil's Araripe Basin. It is an important Lagerstätten, Lagerstätte (undisturbed fossil accumulation) for palaeontology, pa ...
, Brazil **''An. myrmicopterus'' - Aptian Baissa,
Zaza Formation The Zaza Formation is a geological formation located in Buryatia (Russia). It dates to the Lower Cretaceous. The age of the formation is disputed, and is considered likely to be Valanginian-Hauterivian, or Aptian in age. It comprises sandstones, ...
, Russia **''An. niger'' - Aptian, Turga Formation, Russia **''An. pallidus'' - Aptian Gurvan-Eren Formation, Mongolia **''An. parvus'' - Barremian
Crato Formation The Crato Formation is a geologic formation (stratigraphy), formation of Early Cretaceous (Aptian) age in northeastern Brazil's Araripe Basin. It is an important Lagerstätten, Lagerstätte (undisturbed fossil accumulation) for palaeontology, pa ...
, Brazil **''An. penyalveri'' -
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
/
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
La Pedrera de Rúbies Formation, Spain **''An. ryonsangiensis'' - Barremian/Aptian, Sinuiju Formation, North Korea **''An. saxosus'' -
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
, Huangbanjigou,
Yixian Formation The Yixian Formation (; formerly Romanization of Chinese, transcribed as Yihsien Formation or Yixiang Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the Barremian stage of the Early Cretaceous. I ...
, China **''An. strigosus'' -
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
Laiyang Formation, China **''An. venulosus'' -
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
Laiyang Formation, China *'' Archisphex'' (synonyms ''Cretosphex'' ) **''Ar. boothi'' **''Ar. catalunicus'' **''Ar. crowsoni'' **''Ar. curvus'' **''Ar. incertus'' **''Ar. proximus'' *'' Baissodes'' Weald Clay, United Kingdom, Hauterivian Yixian Formation, China, Aptian Gurvan-Eren Formation, Mongolia, Aptian Zaza Formation, Russia, Aptian **''B. grabaui'' **''B. longus'' **''B. magnus'' **''B. robustus'' *'' Cretobestiola'' (synonym ''Bestiola'' ) La Pedrera de Rúbies Formation, Spain, Barremian Crato Formation, Brazil, Aptian, Dzun-Bain Formation, Mongolia, Aptian Zaza Formation, Argun Formation, Russia, Aptian **''Cb. communis'' **''Cb. hispanica'' **''Cb. subpetiolata'' **''Cb. tenuipes'' * '' Cretosphecium'' **''Cs. lobatum'' - Aptian, Dzun-Bain Formation, Mongolia **''Cs. triste'' - Aptian, Dzun-Bain Formation, Mongolia *'' Eosphecium'' **''Eo. naumanni'' -
Ypresian In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by th ...
,
Coldwater Beds The Coldwater Beds are a Formation (geology), geologic formation of the Okanagan Highlands in British Columbia, Canada. They preserve fossils dating back to the Ypresian stage of the Eocene Period (geology), period, or Wasatchian in the North Ame ...
,
Eocene Okanagan Highlands The Eocene Okanagan Highlands or Eocene Okanogan Highlands are a series of Early Eocene geological formations which span a transect of British Columbia, Canada, and Washington state, United States. Known for a highly diverse and detailed plant ...
, Canada *'' Eubaissodes'' **''Eu. completus'' - Aptian, Laiyang Formation, China *'' Ilerdosphex'' **''I. wenzae'' - Barremian, La Pedrera de Rúbies Formation, Spain *'' Mesorhopalosoma'' **''Me. cearae'' - Aptian, Crato Formation, Brazil *'' Montsecosphex'' **''Mo. jarzembowskii'' - Barremian, La Pedrera de Rúbies Formation, Spain *'' Oryctobaissodes'' **''O. armatus'' - Aptian, Zaza Formation, Russia, *'' Paleorhopalosoma'' **''Pa. menatensis'' -
Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
,
Menat Formation The Menat Formation is a geologic formation in France. It preserves fossils dating back to the Paleocene. It is a maar deposit located on top of an ancient volcano, the extent is very localised with the outcropping area being around 600 by 1000 me ...
, France *'' Pompilopterus'' Weald Clay, United Kingdom, Hauterivian/Barremian La Pedrera de Rúbies Formation, Spain, Barremian, Zaza Formation, Russia, Aptian, Sinuiju Formation, North Korea, Aptian **''Po. ciliatus'' **''Po. corpus'' **''Po. difficilis'' **''Po. keymerensis'' **''Po. leei'' **''Po. montsecensis'' **''Po. noguerensis'' **''Po. ryonsangensis'' **''Po. sinuijuensis'' **''Po. wimbledoni'' **''Po. worssami'' *'' Trichobaissodes'' **''T. antennatus'' - Aptian, Zaza Formation, Russia *'' Vitimosphex'' **''V. incompletus'' - Aptian, Zaza Formation, Russia **''V. vividus'' - Yixian Formation, China Undescribed specimens are known from the Cenomanian aged Burmese amber of Myanmar, and several of the Eocene Okanagan Highlands sites. Fossils identified as ''Eosphecium'' sp. have been reported from the Driftwood shales, Tranquille Formation, and
Klondike Mountain Formation The Klondike Mountain Formation is an Early Eocene (Ypresian) geological formation located in the northeast central area of Washington (state), Washington state. The formation is composed of volcanic rocks in the upper unit and volcanic plus lacu ...
. Additional angarosphecid specimens are reported from the Tranquille and
Republic A republic, based on the Latin phrase ''res publica'' ('public affair' or 'people's affair'), is a State (polity), state in which Power (social and political), political power rests with the public (people), typically through their Representat ...
sites plus the
Allenby Formation The Allenby formation is a sedimentary rock formation in British Columbia which was deposited during the Ypresian stage of the Early Eocene. It consists of conglomerates, sandstones with interbedded shales and coal. The shales contain an ...
that are not 'Eosphecium''. '' Burmasphex'' Melo and Rosa (2018) and '' Decasphex'' Zheng ''et al'' (2020), originally assigned to this family, were subsequently moved to the separate apoid family Burmasphecidae.


References

{{Taxonbar, from=Q2849255 Apoidea Prehistoric insect families