Allographa Nana

''Allographa'' is a genus of script lichens in the family Graphidaceae. It has nearly 200 species. Formally circumscribed in 1824 by François Fulgis Chevallier, ''Allographa'' was formerly included in '' Graphis'', but was upgraded to generic status in 2018 by lichenologists Klaus Kalb and Robert Lücking, who used molecular phylogenetics analysis to show this group of species to constitute a distinct lineage in the Graphidaceae.

Description

''Allographa'' is characterised by ascomata (fruiting bodies), which are elongated and groove-like in shape. One of its defining features is the generally excipulum, which gives a dark appearance to the fruiting bodies, although some species formerly placed in ''Hemithecium'' with uncarbonised (blackened) are also included in this genus.

''Allographa'' typically displays robust and strongly prominent lirellae, often with a massive structure. The are hyaline (colourless) and usually large to very large in size, turning violet-blue when treated with iodine. A key distinguishing feature from ''Graphis'' in the strict sense is the strongly gelatinous with thin paraphyses.

While ''Allographa'' shares many morphological similarities with ''Graphis'', making clear-cut distinctions challenging, molecular evidence supports its status as a separate genus. ''Allographa'' is part of a larger clade within Graphidaceae that includes other lirellate genera such as ''Diorygma'', '' Glyphis'', '' Pallidogramme'', ''Phaeographis'', ''Platygramme'', ''Platythecium'', ''Sarcographa'', ''Thalloloma'', and ''Thecaria''.

The genus encompasses species previously assigned to the ''Graphis carassensis'' group and some former ''Hemithecium'' species. Notable examples include '' A. acharii'', '' A. chrysocarpa'', '' A. rhizocola'', '' A. ruiziana'', and species related to '' A. chlorocarpa'' and '' A. rufopallida''. Despite the morphological overlap with ''Graphis'', advanced techniques such as morphology-based phylogenetic binning have successfully differentiated most ''Allographa'' species from ''Graphis'' sensu stricto, confirming its status as a distinct genus within the Graphidaceae.

Species

*''Allographa acharii''
*'' Allographa adpressa''
*'' Allographa albotecta''
*'' Allographa altamirensis''
*'' Allographa amazonica''
*'' Allographa ancelica'' – Brazil
*'' Allographa anguilliradians''
*'' Allographa angustata''
*'' Allographa apertoinspersa''
*'' Allographa apicalinspersa'' – Brazil
*'' Allographa apoda''
*'' Allographa aptrootiana''
*'' Allographa aquilonia''
*''Allographa argentata''
*'' Allographa asterizans''
*'' Allographa atrocelata''
*'' Allographa atrocelatoides''
*'' Allographa balbisii''
*'' Allographa balbisina''
*'' Allographa bambusicola'' – Sri Lanka
*'' Allographa barillasiae''
*'' Allographa bettinae''
*'' Allographa bifera''
*'' Allographa brachylirellata''
*'' Allographa bukittimaensis''
*'' Allographa calcea''
*'' Allographa cameroonensis''
*'' Allographa carassensis''
*'' Allographa celata''
*'' Allographa cerradensis''
*'' Allographa chamelensis''
*'' Allographa chloroalba''
*''Allographa chlorocarpa''
*''Allographa chrysocarpa''
*'' Allographa cinerea''
*'' Allographa cleistoblephara''
*'' Allographa cleistomma''
*'' Allographa cognata''
*'' Allographa comma''
*'' Allographa congesta''
*'' Allographa conglomerata''
*'' Allographa consanguinea''
*'' Allographa contorta''
*'' Allographa contortuplicata''
*'' Allographa curtiuscula''
*'' Allographa cylindrospora''
*'' Allographa daintreensis''
*'' Allographa dealbata''
*'' Allographa dispersa''
*'' Allographa dolichographa''
*'' Allographa dotalugalensis''
*'' Allographa elevativerrucosa''
*'' Allographa elixii''
*'' Allographa elmeri''
*'' Allographa elongata''
*'' Allographa epixantha''
*'' Allographa exuens''
*'' Allographa exuta''
*'' Allographa farinulenta''
*'' Allographa firferi''
*'' Allographa flabillata''
*'' Allographa flavens''
*'' Allographa flavoaltamirensis''
*'' Allographa flavominiata''
*'' Allographa fournieri''
*'' Allographa frumentaria''
*'' Allographa fujianensis''
*'' Allographa funilina''
*'' Allographa glauconigra''
*'' Allographa globosa''
*'' Allographa gomezii''
*'' Allographa gracilescens''
*'' Allographa grandis''
*'' Allographa granulata''
*''Allographa granulosa''
*'' Allographa gregmuelleri''
*'' Allographa guainiae''
*'' Allographa hadrospora''
*'' Allographa hainanense''
*'' Allographa hossei''
*'' Allographa hypostictica''
*'' Allographa illinata''
*'' Allographa immersa''
*'' Allographa incerta''
*'' Allographa ingarum''
*'' Allographa inspersostictica''
*''Allographa inturgescens''
*'' Allographa invisibilis''
*'' Allographa isidiata''
*'' Allographa itatiaiensis''
*'' Allographa jayatilakana'' – Sri Lanka
*'' Allographa kamojangensis''
*'' Allographa kansriana''
*'' Allographa knucklensis''
*'' Allographa kuetchangiana''
*'' Allographa labiata''
*'' Allographa laubertiana''
*''Allographa lecanactiformis''
*'' Allographa leprographa''
*'' Allographa leptospora''

*''Allographa leucaenae''
*'' Allographa lichexanthonica''
*'' Allographa longilirellata''
*'' Allographa longissima''
*'' Allographa longula''
*''Allographa lourdesina''
*'' Allographa lumbricina''
*'' Allographa lumbschii''
*'' Allographa lutea''
*'' Allographa macella''
*'' Allographa mahaeliyensis''
*'' Allographa malacodes''
*'' Allographa marginata''
*''Allographa medioinspersa'' – Brazil
*'' Allographa mexicana''
*'' Allographa miniata''
*'' Allographa mirabilis''
*'' Allographa multistriata''
*'' Allographa multisulcata''
*'' Allographa myolensis''
*'' Allographa nadurina''
*'' Allographa nana''
*'' Allographa norlabiata''
*'' Allographa norvestitoides''
*'' Allographa nuda''
*'' Allographa nudaeformis''
*'' Allographa obtectostriata''
*'' Allographa ochracea''
*'' Allographa olivacea''
*'' Allographa oryzaecarpa''
*'' Allographa oryzaeformis''
*'' Allographa ovata''
*'' Allographa oxyspora''
*''Allographa pachygrapha''
*'' Allographa parallela''
*'' Allographa patwardhanii''
*'' Allographa pavoniana''
*''Allographa pedunculata''
*'' Allographa phaeospora''
*'' Allographa pilarensis''
*'' Allographa pitmanii''
*'' Allographa pittieri''
*'' Allographa plagiocarpa''
*''Allographa plumbea''
*'' Allographa plurispora''
*'' Allographa polillensis''
*'' Allographa polystriata''
*'' Allographa pseudoaquilonia''
*'' Allographa pseudocinerea''
*'' Allographa pruinodisca'' – Brazil
*'' Allographa regularis''
*'' Allographa rhizicola''
*'' Allographa rimulosa''
*'' Allographa rufopallida''
*''Allographa ruiziana''
*'' Allographa rustica''
*'' Allographa salacinicum''
*''Allographa salacinilabiata''
*'' Allographa sarawakensis''
*'' Allographa sauroidea''
*'' Allographa sayeri''
*'' Allographa scaphella''
*'' Allographa schummii''
*'' Allographa seminuda''
*'' Allographa semirigida''
*'' Allographa sessilis''
*'' Allographa singaporensis''
*'' Allographa sitiana''
*'' Allographa sitianoides''
*'' Allographa sorsogona''
*'' Allographa sterlingiana''
*'' Allographa stictilabiata''
*'' Allographa striatula''
*'' Allographa subamylacea''
*'' Allographa subassimilis''
*'' Allographa subcelata''
*'' Allographa subdisserpens''
*'' Allographa subdussii''
*'' Allographa subelmeri''
*'' Allographa subflexibilis''
*''Allographa subimmersa''
*'' Allographa submirabilis''
*'' Allographa subradiata''
*'' Allographa suprainspersata''
*'' Allographa subruiziana''
*'' Allographa subserpens''
*'' Allographa subturgidula''
*'' Allographa superans''
*'' Allographa supertecta''
*'' Allographa tamiamiensis''
*'' Allographa trichospora''
*'' Allographa triphora''
*'' Allographa tumidula''
*'' Allographa upduna''
*'' Allographa upretii''
*'' Allographa uruguayensis''
*'' Allographa vandenboomiana''
*'' Allographa verminosa''
*'' Allographa vernicosa''
*'' Allographa vestita''
*'' Allographa vestitoides''
*'' Allographa wangii''
*'' Allographa weerasooriyana'' – Sri Lanka
*'' Allographa weii''
*''Allographa xanthospora''

References

{{Taxonbar, from=Q104721240

Lichen genera
Graphidales genera
Taxa described in 1824
Taxa named by François Fulgis Chevallier