Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian

geologist A geologist is a scientist who studies the structure, composition, and History of Earth, history of Earth. Geologists incorporate techniques from physics, chemistry, biology, mathematics, and geography to perform research in the Field research, ...

and

paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...

who is a leading expert in the field of studying

pterosaurs Pterosaurs are an extinct clade of flying reptiles in the Order (biology), order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosau ...

. His research has focused mainly on

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

reptiles from the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...

Period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...

, including extinct

dinosaurs Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...

and

crocodylomorphs Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. Extinct crocodylomorphs were considerably more ...

. Kellner has over 500 publications to his name, has published more than 160 primary studies and two science books. He has participated in paleontological expeditions to many locations including Brazil, Chile, Iran, the United States, Argentina, China, and Antarctica. His scientific achievements include the description of more than thirty

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

. For his work he has received several honors and prizes, including the

TWAS Prize This is a list of recipients of the TWAS Prize, awarded annually by The World Academy of Sciences (TWAS) since 1985. Summary Agricultural Sciences Biology Chemistry Earth Sciences Engineering Sciences Mathematics ...

for Earth Sciences from

The World Academy of Sciences The World Academy of Sciences for the advancement of science in developing countries (TWAS) is a North–south research partnerships, merit-based science academy established for developing countries, uniting more than 1,400 scientists in some 1 ...

and admission to the

National Order of Scientific Merit The National Order of Scientific Merit () is an honor bestowed upon Brazilian and foreign personalities recognized for their scientific and technical contributions to the cause and development of science in Brazil. It was instituted on March 16, 19 ...

(class Comendador), one Brazil's most prestigious awards.

Biography

Kellner was born in

Vaduz Vaduz (; or ; High Alemannic pronunciation: [])Hans Stricker, Toni Banzer, Herbert Hilbe: ''Liechtensteiner Namenbuch. Die Orts- und Flurnamen des Fürstentums Liechtenstein.'' Band 2: ''Die Namen der Gemeinden Triesenberg, Vaduz, Schaan.'' ...

Liechtenstein Liechtenstein (, ; ; ), officially the Principality of Liechtenstein ( ), is a Landlocked country#Doubly landlocked, doubly landlocked Swiss Standard German, German-speaking microstate in the Central European Alps, between Austria in the east ...

, son of a German father and Austrian mother. In his early childhood he moved with his parents to Brazil, where he became naturalised Brazilian. In

Rio de Janeiro Rio de Janeiro, or simply Rio, is the capital of the Rio de Janeiro (state), state of Rio de Janeiro. It is the List of cities in Brazil by population, second-most-populous city in Brazil (after São Paulo) and the Largest cities in the America ...

he received a

primary Primary or primaries may refer to: Arts, entertainment, and media Music Groups and labels * Primary (band), from Australia * Primary (musician), hip hop musician and record producer from South Korea * Primary Music, Israeli record label Work ...

and

secondary education Secondary education is the education level following primary education and preceding tertiary education. Level 2 or ''lower secondary education'' (less commonly ''junior secondary education'') is considered the second and final phase of basic e ...

at the bilingual Escola Alemã Corcovado. He began studying

geology Geology (). is a branch of natural science concerned with the Earth and other astronomical objects, the rocks of which they are composed, and the processes by which they change over time. Modern geology significantly overlaps all other Earth ...

at the

Federal University of Rio de Janeiro The Federal University of Rio de Janeiro (, UFRJ) is a public university, public research university in Rio de Janeiro, Brazil. It is the largest federal university in the country and is one of the Brazilian centers of excellence in teaching and r ...

(UFRJ) in 1981. As a student, he soon began researching

vertebrates Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebra ...

, particularly

reptiles Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...

from Brazil's

. His studies largely concerned

pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...

specimens from the geologic

Santana Group The Santana Group is a geologic group, formerly included as the middle part of the Araripe Group, in the Araripe Basin of northeastern Brazil. The group comprises the Crato, Ipubi and Romualdo Formations and is dated to the Aptian to Albian stag ...

, about which he published many papers in the late 1980s. He earned a

Master of Science A Master of Science (; abbreviated MS, M.S., MSc, M.Sc., SM, S.M., ScM or Sc.M.) is a master's degree. In contrast to the Master of Arts degree, the Master of Science degree is typically granted for studies in sciences, engineering and medici ...

degree in geology at the UFRJ in 1991, a

MPhil A Master of Philosophy (MPhil or PhM; Latin ' or ') is a postgraduate degree. The name of the degree is most often abbreviated MPhil (or, at times, as PhM in other countries). MPhil are awarded to postgraduate students after completing at least ...

degree in 1994, and a

Ph.D. A Doctor of Philosophy (PhD, DPhil; or ) is a terminal degree that usually denotes the highest level of academic achievement in a given discipline and is awarded following a course of graduate study and original research. The name of the deg ...

in 1996 from

Columbia University Columbia University in the City of New York, commonly referred to as Columbia University, is a Private university, private Ivy League research university in New York City. Established in 1754 as King's College on the grounds of Trinity Churc ...

in New York, in a joint program with the

American Museum of Natural History The American Museum of Natural History (AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. Located in Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 21 interconn ...

. In 1997 he became a professor at the Federal University-owned

National Museum of Rio de Janeiro National may refer to: Common uses * Nation or country ** Nationality – a ''national'' is a person who is subject to a nation, regardless of whether the person has full rights as a citizen Places in the United States * National, Maryland, ce ...

and

curator A curator (from , meaning 'to take care') is a manager or overseer. When working with cultural organizations, a curator is typically a "collections curator" or an "exhibitions curator", and has multifaceted tasks dependent on the particular ins ...

of its geological and

paleontological Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...

departments. From 1998 to 2001, he served as the chairman of these departments and has been head of the

zoology Zoology ( , ) is the scientific study of animals. Its studies include the anatomy, structure, embryology, Biological classification, classification, Ethology, habits, and distribution of all animals, both living and extinction, extinct, and ...

graduate program since 2008. Kellner is also chief editor of the '' Anais da Academia Brasileira de Ciências'', the official publication of the

Brazilian Academy of Sciences Brazilian commonly refers to: * Brazil, a country * Brazilians, its people * Brazilian Portuguese, its dialect Brazilian may also refer to: * "The Brazilian", a 1986 instrumental music piece by Genesis * Brazilian Café, Baghdad, Iraq (1937) * Bra ...

Alexander_Kellner_at_the_Academia_Brasileira_de_Sciencias

As part of his work on flying reptiles, Kellner organised the Pterosaur Workshop at Pittsburgh, USA, in 1995, and the first pterosaur

symposium In Ancient Greece, the symposium (, ''sympósion'', from συμπίνειν, ''sympínein'', 'to drink together') was the part of a banquet that took place after the meal, when drinking for pleasure was accompanied by music, dancing, recitals, o ...

ever held at the American Museum of Natural History in 1996. As well, he was involved in the organisation of several scientific meetings in Rio de Janeiro such as the 31st International Geological Congress in 2000 and the 2nd Latin-American Congress of Vertebrate Paleontology in 2002. Kellner has organized or taken part in several paleontological expeditions to many locations around the globe, including Brazil—

Mato Grosso Mato Grosso ( – ) is one of the states of Brazil, the List of Brazilian states by area, third largest by area, located in the Central-West Region, Brazil, Central-West region. The state has 1.66% of the Brazilian population and is responsible ...

Rio Grande do Sul Rio Grande do Sul (, ; ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative units of Brazil#List, fifth-most populous state and the List of Brazilian s ...

and

Ceará Ceará (, ) is one of the 26 states of Brazil, located in the Northeast Region, Brazil, northeastern part of the country, on the Atlantic Ocean, Atlantic coast. It is the List of Brazilian states by population, eighth-largest Brazilian State by ...

; the deserts of

Atacama The Atacama Desert () is a desert plateau located on the Pacific coast of South America, in the north of Chile. Stretching over a strip of land west of the Andes Mountains, it covers an area of , which increases to if the barren lower slopes ...

, Chile and

Kerman Kerman (; ) is a city in the Central District (Kerman County), Central District of Kerman County, Kerman province, Kerman province, Iran, serving as capital of the province, the county, and the district. History Kerman was founded as a def ...

, Iran;

Montana Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Idaho to the west, North Dakota to the east, South Dakota to the southeast, Wyoming to the south, an ...

in the United States;

Patagonia Patagonia () is a geographical region that includes parts of Argentina and Chile at the southern end of South America. The region includes the southern section of the Andes mountain chain with lakes, fjords, temperate rainforests, and glaciers ...

in Argentina; the famous deposits of

Liaoning ) , image_skyline = , image_alt = , image_caption = Clockwise: Mukden Palace in Shenyang, Xinghai Square in Dalian, Dalian coast, Yalu River at Dandong , image_map = Liaoning in China (+all claims hatched).svg , ...

, China, and

James Ross Island James Ross Island () is a large island off the southeast side and near the northeastern extremity of the Antarctic Peninsula, from which it is separated by Prince Gustav Channel. Rising to , it is irregularly shaped and extends in a north–so ...

in Antarctica. Kellner has over 500 publications to his name (including abstracts and science articles). He has published more than 160 primary studies and two science books: ''Pterossauros - os senhores do céu do Brasil'' ("Pterosaurs — Lords of the Brazilian Sky") and the novel ''Na terra dos titãs'' ("In the Land of the Titans"). He has also taken part in documentaries about fossils (e.g., ''Antarctica - a Summer of 70 million Years'' and ''Dinosaur Hunters''). As a result of his scientific activity he has received several honours, being appointed a member of the Brazilian Academy of Sciences in 1997. He is also an honourable member of the New York Paleontologial Society and the Paleontological Society of Chile. He is a research associate of the American Museum of Natural History and of the Chinese

Institute of Vertebrate Paleontology and Paleoanthropology The Institute of Vertebrate Paleontology and Paleoanthropology (IVPP; ) of People's Republic of China, China is a research institution and collections repository for fossils, including many dinosaur and pterosaur specimens (many from the Yixian For ...

. Besides his teaching activity, having advised over fifteen master and Ph.D. students, Kellner has been active in the propagation of scientific knowledge to the general public. He organised the 1999 exposition ''No Tempo dos Dinossauros'' ("In the Time of the Dinosaurs") at the Museum of Earth Sciences, which has been regarded as a landmark for the establishment of paleontology in Brazil, attracting the attention of the people of Brazilian to the studies of fossils. In 2006 he organized the mounting of the first large-scale dinosaur skeleton in Brazil, that of the

sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...

'' Maxakalisaurus topai'', for which he received recognition from the

Brazilian Congress The National Congress () is the legislative body of Brazil's federal government. Unlike the state legislative assemblies and municipal chambers, the Congress is bicameral, composed of the Federal Senate (the upper house) and the Chamber of D ...

. Since 2004 he has written a monthly column in ''Caçadores de Fósseis'' (Fossil Hunters), on the website ''Ciência Hoje On Line'', a project of the Brazilian Society for Scientific Progress. Apart from studying their fossils, Kellner has performed important theoretical work on pterosaurs, including

cladistic Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...

studies regarding their

phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ...

. In this he is the founder of a distinctive Brazilian school of the study of pterosaurs, with its own favoured phylogenetic model, clade terminology and nomenclature. Rival models and nomenclatural choices have been devised by the influential British pterosaur researcher

David Unwin David (; , "beloved one") was a king of ancient Israel and Judah and the third king of the United Monarchy, according to the Hebrew Bible and Old Testament. The Tel Dan stele, an Aramaic-inscribed stone erected by a king of Aram-Damas ...

. Kellner has received numerous honours and prizes, including the

for Earth Sciences from

. He was admitted in the

(class Comendador), one the Brazilian Federal Government's most prestigious awards.

List of species named by Kellner

Kellner's scientific achievements include the description of more than thirty

, of which ''

Santanaraptor ''Santanaraptor'' (meaning " Santana Formation thief") is a genus of tyrannosauroid theropod dinosaur that lived in South America during the Early Cretaceous (late Aptian-early Albian), about 112 million years ago. Discovery The type species i ...

'' ''placidus'' (1996, 1999) is among the best examples of soft tissue preservation, including blood vessels and muscle fibers, reported in any

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...

. The pterosaur ''

Thalassodromeus ''Thalassodromeus'' is a genus of pterosaur that lived in what is now Brazil during the Early Cretaceous Period (geology), period, about a hundred million years ago. The original skull, discovered in 1983 in the Araripe Basin of northeastern B ...

sethi'', which Kellner described in 2002 with his colleague Diogenes de Almeida Campos, allowed for the establishment of a new hypothesis regarding the use of the head crest in body temperature regulation of pterosaurs. A complete list of new species described and named by Kellner, sometimes in cooperation with other researchers, includes: 20130825_Brazil_Rio_de_Janeiro_0144

*''

Brasileodactylus ''Brasileodactylus'' a genus of pterosaur from the Aptian-age (Early Cretaceous period) lower Santana formation of Chapada do Araripe, Ceará, Brazil. The genus was named by paleontologist Alexander Wilhelm Armin Kellner in 1984. The genus na ...

araripensis'' Kellner, 1984 (

Reptilia Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and amniotic development. Living traditional reptiles comprise four orders: Testudines, Crocodilia, Squamata, and Rhynchocephalia. About 12,000 living spe ...

, Pterosauria) * ''

Anhanguera Anhanguera may refer to: People * Bartolomeu Bueno da Silva (1672–1740), a bandeirante also known as Anhanguera * Anhangüera I, father of Bartolomeu Bueno da Silva Places in Brazil * Anhanguera, Goiás, a municipality in the state of Goiás ...

blittersdorffi'' Campos & Kellner, 1985 (Reptilia, Pterosauria) * '' Oshunia brevis'' Wenz & Kellner, 1986 (

Pisces Pisces may refer to: *Pisces (astrology), an astrological sign Astronomy *Pisces (constellation), a constellation ** Pisces Overdensity, an overdensity of stars in the Milky Way's halo that is situated in the Pisces constellation ** Pisces II, a ...

Halecomorphi Halecomorphi is a taxon of Actinopterygii, ray-finned bony fish in the clade Neopterygii. The only extant Halecomorph species are the bowfin (''Amia calva'') and eyespot bowfin (''Amia ocellicauda''), but the group contains many extinct species i ...

). * ''

Caririsuchus ''Caririsuchus'' is an extinct genus of peirosaurid crocodylomorph. Fossils have been found from the Romualdo Formation of the Santana Group in the Araripe Basin in northeastern Brazil, dating back to the Albian stage of the Early Cretaceous Th ...

camposi'' Kellner, 1987 (Reptilia,

Crocodylia Crocodilia () is an Order (biology), order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorp ...

) * ''

Tupuxuara ''Tupuxuara'' is a genus of thalassodromid pterosaur that lived during the Albian age of the Early Cretaceous, about 112 million years ago. Its remains were found in what is now the Romualdo Formation of the Santana Group in Brazil. ''Tupuxuara ...

longicristatus'' Kellner & Campos, 1988 (Reptilia, Pterosauria) * '' Tapejara wellnhoferi'' Kellner, 1989 (Reptilia, Pterosauria) * ''Tupuxuara leonardii'' Kellner & Campos, 1994 (Reptilia, Pterosauria) * '' Angaturama limai'' Kellner & Campos, 1996 (Reptilia, Dinosauria) * '' Ongghonia dashzevegi'' Kellner & McKenna, 1996 (

Mammalia A mammal () is a vertebrate animal of the class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three middle ear bon ...

, Leptictidae) * ''

Tupandactylus ''Tupandactylus'' (meaning "Tupan finger", in reference to the Tupi thunder god) is a genus of tapejarid pterodactyloid pterosaur from the Early Cretaceous Crato Formation of Brazil. History ''Tupandactylus imperator'' is known from four nea ...

imperator'' (Campos & Kellner, 1997) (Reptilia, Pterosauria) * ''

Siroccopteryx ''Siroccopteryx'' is an extinct genus of anhanguerid pterodactyloid pterosaur which lived in Morocco during the Cenomanian stage of the Late Cretaceous. Some researchers, such as David M. Unwin, consider the genus a junior synonym of '' Colobor ...

moroccensis'' Mader & Kellner, 1999 (Reptilia, Pterosauria) * ''

Gondwanatitan ''Gondwanatitan'' (meaning "giant from Gondwana") was a titanosaurian sauropod dinosaur. ''Gondwanatitan'' was found in Brazil, at the time part of the southern supercontinent Gondwana, in the late Cretaceous Period (70 mya). Like some other sau ...

faustoi'' Kellner & Azevedo, 1999 (Reptilia, Dinosauria) * ''

placidus'' Kellner, 1999 (Reptilia, Dinosauria) * ''Anhanguera piscator'' Kellner & Tomida, 2000 (Reptilia, Pterosauria) * ''

Stratiotosuchus ''Stratiotosuchus'' (from Greek, (stratiōtēs, "soldier") and (suchos, "crocodile")) is an extinct genus of baurusuchid mesoeucrocodylian from the Adamantina Formation in Brazil. It lived during the Late Cretaceous. The first fossils were foun ...

maxhechti'' Campos, Suarez, Riff & Kellner, 2001 (Reptilia, Crocodylia) * ''

sethi'' Kellner & Campos, 2002 (Reptilia, Pterosauria) Thalassodromeus in Japan

* '' Pycnonemosaurus nevesi'' Kellner & Campos, 2002 (Reptilia, Dinosauria) * ''

Kaikaifilusaurus ''Kaikaifilusaurus'' is an extinct genus of rhynchocephalians in the family Sphenodontidae from the Late Cretaceous of South America. Fossils of the genus were found in Cenomanian sediments of the Candeleros Formation and Turonian layers of the ...

calvoi'' Simón & Kellner, 2003. (Reptilia, Sphenodontia) * ''

Unaysaurus ''Unaysaurus'' is a genus of unaysauridae, unaysaurid sauropodomorpha, sauropodomorph herbivore dinosaur. Discovered in southern Brazil, in the geopark of Paleorrota, in 1998, and announced in a press conference on Thursday, December 3, 2004, it ...

tolentinoi'' Leal, L.A., Azevedo, S.A., Kellner, A.W.A. & Rosa, Á.A.S., 2004 (Reptilia, Dinosauria) * ''

Unenlagia ''Unenlagia'' (meaning "half-bird" in Latinized Mapudungun) is a genus of dromaeosaurid theropod dinosaur that lived in South America during the Late Cretaceous period. The genus ''Unenlagia'' has been assigned two species: ''U. comahuensis'', ...

paynemili'' Calvo, J.O., Porfiri, J. & Kellner, A.W.A., 2004 (Reptilia, Dinosauria) * ''

Feilongus ''Feilongus'' is an extinct genus of ctenochasmatid pterodactyloid pterosaur from the Barremian–Aptian-age Lower Cretaceous Yixian Formation of Beipiao, Liaoning, China. Discovery and naming The genus was named and described in 2005 by Wang X ...

youngi'' Wang, Kellner, Zhou & Campos, 2005 (Reptilia, Pterosauria) * ''

Nurhachius ''Nurhachius'' is a genus of Istiodactylidae, istiodactylid pterodactyloid pterosaur from the Barremian to Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, Chaoyang, Liaoning, China. Its fossil remains date back about 120 mi ...

ignaciobritoi'' Wang, Kellner, Zhou & Campos, 2005 (Reptilia, Pterosauria) * ''

Baurutitan ''Baurutitan'' is a genus of sauropod dinosaur that lived during the Late Cretaceous in what is now Brazil. The type species, ''Baurutitan britoi'', was described in 2005 by Kellner and colleagues, although the fossil remains had already been di ...

britoi'' Kellner, A.W.A., Campos, D. A., Trotta, M. N. F. 2005 (Reptilia, Dinosauria) * '' Trigonosaurus pricei'' Campos, D.A., Kellner, A.W.A., Bertini, R.J., Santucci, R.M. 2005 (Reptilia, Dinosauria) * '' Caririemys violetae'' Oliveira, G.R. & Kellner, A.W.A. 2007 (Reptilia, Testudines) * ''

Gegepterus ''Gegepterus'' is a genus of ctenochasmatid pterosaur from the Early Cretaceous period of what is now the Yixian Formation of Liaoning, China. Only one species is known, ''G. changi''. History and etymology The genus was named in 2007 by Wang Xi ...

changi'' Wang, Kellner, Zhou & Campos, 2007 (Reptilia, Pterosauria) * ''

Futalognkosaurus ''Futalognkosaurus'' ( ; meaning "giant chief lizard") is a genus of titanosaurian dinosaur. The herbivorous ''Futalognkosaurus'' lived approximately 87 million years ago in the Portezuelo Formation, in what is now Argentina, of the Coniacian st ...

dukei'' Calvo, J.O., Porfiri, J., González-Riga, B.J. & Kellner, A.W.A., 2007 (Reptilia, Dinosauria) * '' Nemicolopterus crypticus'' Wang, Kellner, Zhou & Campos, 2008 (Reptilia, Pterosauria) * '' Guarinisuchus munizi'' Barbosa, Kellner & Viana, 2008 (Reptilia, Crocodylia) * ''

Hongshanopterus ''Hongshanopterus'' is a genus of pterodactyloid pterosaur from the Lower Cretaceous Jiufotang Formation of Liaoning, China. The type species ''Hongshanopterus lacustris'' was in 2008 named and described by Wang Xiaolin, Diogenes de Almeida C ...

lacustris'' Wang, Zhou, Campos & Kellner, 2008 (Reptilia, Pterosauria) * ''

Coringasuchus ''Coringasuchus'' is a genus of mesoeucrocodylian crocodyliform, perhaps a notosuchian. It is known from fossils including cranial material discovered in rocks of the early Cenomanian-age Upper Cretaceous Alcântara Formation, Cajual Island, n ...

anisodontis'' Kellner, A.W.A., Pinheiro, A.E.P., Azevedo, S.A.K., Henriques, D.D.R., Carvalho, L.B. & Oliveira, G. 2009 (Reptilia, Crocodylia) * ''

Wukongopterus ''Wukongopterus'' is a genus of basal pterosaur, found in Liaoning, China, from the Tiaojishan Formation, of the Middle or Late Jurassic. It was unusual for having both an elongate neck and a long tail. Discovery and naming The genus was desc ...

lii'' Wang, Kellner, Jiang, Meng 2009 (Reptilia, Pterosauria) * ''

Dawndraco ''Dawndraco'' is a controversial genus of pteranodontidae, pteranodontid pterosaur from the Late Cretaceous of North America. It is likely synonymous with the contemporary pteranodontid ''Geosternbergia.'' ''Dawndraco'' was named by Alexander K ...

kanzai'', Kellner 2010 (Reptilia, Pterosauria) * ''

Geosternbergia ''Pteranodon sternbergi'' is an extinct species of the pteranodontid pterodactyloid pterosaur genus ''Pteranodon'' from the Late Cretaceous geological period of North America. ''P. sternbergi'' was among the largest pterosaurs, with a wingspan ...

maysei'', Kellner 2010 (Reptilia, Pterosauria) * ''

Aussiedraco ''Aussiedraco'' is a genus of targaryendraconian pterodactyloid pterosaur from the early Cretaceous of Australia. Description ''Aussiedraco'' is known from holotype QM F10613, a partial mandibular symphysis housed at the Queensland Museum ...

molnari'', Kellner, A.W.A., Rodrigues, T., Costa, F.R., 2011 (Reptilia, Pterosauria) * ''

Oxalaia ''Oxalaia'' (in reference to the African deity ''Oxalá'') is a controversial genus of Spinosauridae, spinosaurid dinosaur that lived in what is now the Northeast Region, Brazil, Northeast Region of Brazil during the Cenomanian Geological stage, ...

quilombensis'', Kellner, A.W.A., Machado E.B., Azevedo, S.A.K., Henriques D.R., Carvalho, L.B., 2011 (Reptilia, Dinosauria) * ''

Europejara ''Europejara'' is a genus of tapejarid pterosaur from the Early Cretaceous Period of Spain. The type species, type and only species known is ''Europejara olcadesorum''. Discovery and naming In 2012, the type species ''Europejara olcadesorum'' ...

olcadesorum'', Vullo, Marugán-Lobón, Kellner, et al. 2012 (Reptilia, Pterosauria) * ''

Guidraco ''Guidraco'' (Chin. ''gui'' (鬼) "malicious ghost" + Lat. ''draco'' "dragon") is an extinct genus of toothed pterodactyloid pterosaur known from the Early Cretaceous of Liaoning Province, northeast China. According to many recent studies, ''Gu ...

venator'', Wang, Kellner, et al. 2012 (Reptilia, Pterosauria) * '' Caupedactylus ybaka'', Kellner 2013 (Reptilia, Pterosauria) * ''

Caiuajara ''Caiuajara'' is an extinct genus of tapejarid pterosaur from the Early Cretaceous period (Aptian to Albian stages) of Brazil. It is known from a single type species, ''Caiuajara dobruskii''. Discovery and naming In 1971, the labourers Alexan ...

dobruskii'', Manzig, Kellner, et al. 2014 (Reptilia, Pterosauria) * ''

Hamipterus ''Hamipterus'' is an extinct genus of pteranodontoid pterosaurs from the Early Cretaceous Shengjinkou Formation of northwestern China. It is known from a single species, the type species, ''H. tianshanensis''. Discovery and naming In 2006 from ...

tianshanensis'', Wang, Kellner, et al. 2014 (Reptilia, Pterosauria) * '' Ikrandraco avatar'', Wang, Rodrigues, Jiang, Cheng, Kellner 2014 (Reptilia, Pterosauria) * ''

Austriadraco ''Austriadraco'' is a genus of pterosaur living during the Late Triassic (Norian stage, 217-213 million years ago)Müller R.T., Ezcurra M.D., Garcia M.S., Agnolín F.L., Stocker M.R., Novas F.E., Soares M.B., Kellner A.W.A. & Nesbitt S.J. (2023). ...

dallavecchiai'', Kellner 2015 (Reptilia, Pterosauria) * ''

Bergamodactylus ''Bergamodactylus'' is a putative genus of basal pterosaur which lived during the Late Triassic (early Norian) in the area of present-day Bergamo province in Italy. Its only species is ''Bergamodactylus wildi''. It was named in 2015 based on a p ...

wildi'', Kellner 2015 (Reptilia, Pterosauria) * ''

Linlongopterus ''Linlongopterus'' is a genus of pteranodontoidea, pteranodontoid pterodactyloidea, pterodactyloid pterosaur from the Early Cretaceous of China. It is known from a partial skull and mandible first named and described in 2015 by Rodrigues ''et al. ...

jennyae'', Rodrigues, T., Jiang, S., Cheng, X., Wang, X., & Kellner, A.W.A., 2015 (Reptilia, Pterosauria) * '' Aymberedactylus cearensis'', Pêgas, R.V., Leal, M.E.C., & Kellner, A.W.A., 2016 (Reptilia, Pterosauria) * ''

Argentinadraco ''Argentinadraco'' (meaning "Argentina dragon") is an extinct genus of azhdarchoidea, azhdarchoid pterosaur from the Late Cretaceous Portezuelo Formation of Argentina. It contains a single species, ''A. barrealensis'', named in 2017 by Alexander ...

barrealensis'', Kellner & Calvo, 2017 (Reptilia, Pterosauria) * ''

Iberodactylus ''Iberodactylus'' is a genus of pterodactyloid pterosaurs belonging to the clade Anhangueria, that during the Early Cretaceous lived in the area of present Spain. The type species is ''Iberodactylus andreui''. History of discovery and naming I ...

andreui'', Holgado, B., Pêgas, R.V., Canudo, J.I., Fortuny, J., Rodrigues, T., Company, J., & Kellner, A.W.A., 2019 (Reptilia, Pterosauria) * ''

Keresdrakon ''Keresdrakon'' is an extinct genus of azhdarchoid pterosaur from the Goio-Erê Formation of Brazil, which dates back to the Early Cretaceous period (Aptian-Albian stages), 125 to 100.5 million years ago. ''Keresdrakon'' contains a single spec ...

vilsoni'', Kellner, A.W.A., Weinschütz, L.C., Holgado, B., Bantim, R.A. & Sayão, J.M., 2019 (Reptilia, Pterosauria) * ''

Mimodactylus ''Mimodactylus'' is a genus of istiodactyliform pterosaur that lived in what is now Lebanon during the Late Cretaceous, 95 million years ago. The only known specimen was discovered in a limestone quarry near the town of Hjoula, belonging to ...

libanensis'', Kellner, A.W.A., Caldwell, M.W., Holgado, B., Dalla Vecchia, F.M., Nohra, R., Sayão, J.M. & Currie, P.J., 2019 (Reptilia, Pterosauria)

References

* http://lattes.cnpq.br/0424535851535945 * Pterosaur's Wing, "Hairs" Unlike Any Living Animals, Charles Q. Choi, for National Geographic News (August 4, 2009)

External links

* Museu Nacional/UFRJ - http://acd.ufrj.br/mndgp/kellner/p/index.html {{DEFAULTSORT:Kellner, Alexander 1961 births Living people People from Vaduz Brazilian people of German descent Brazilian people of Austrian descent Brazilian paleontologists TWAS laureates Scientists from Rio de Janeiro (city) Federal University of Rio de Janeiro alumni Columbia University alumni People associated with the American Museum of Natural History National Museum of Brazil