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Varanid
The Varanidae are a family of lizards in the superfamily Varanoidea and order Anguimorpha. The family, a group of carnivorous and frugivorous lizards, includes the living genus '' Varanus'' and a number of extinct genera more closely related to ''Varanus'' than to the earless monitor lizard (''Lanthanotus''). ''Varanus'' includes the Komodo dragon (the largest living lizard), crocodile monitor, savannah monitor, the goannas of Australia and Southeast Asia, and various other species with a similarly distinctive appearance. Their closest living relatives are the earless monitor lizard and Chinese crocodile lizard. The oldest members of the family are known from the Late Cretaceous of Mongolia. Taxonomy The Varanidae were defined (using morphological characteristics) by Estes, de Queiroz and Gauthier (1988) as the clade containing the most recent common ancestor of '' Lanthanotus'' and ''Varanus'' and all of its descendants. A similar definition was formulated by Conrad ' ...
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Komodo Dragon
The Komodo dragon (''Varanus komodoensis''), also known as the Komodo monitor, is a large reptile of the monitor lizard family Varanidae that is endemic to the Indonesian islands of Komodo (island), Komodo, Rinca, Flores, Gili Dasami, and Gili Motang. The largest extant population lives within the Komodo National Park in Eastern Indonesia. It is the List of largest extant lizards, largest extant species of lizard, with the males growing to a maximum length of and weighing up to . As a result of their size, Komodo dragons are apex predators, and dominate the ecosystems in which they live. Komodo dragons hunt and ambush prey including invertebrates, birds, and mammals. Komodo dragons' group behavior in hunting is exceptional in the reptile world. The diet of Komodo dragons mainly consists of Javan rusa (''Rusa timorensis''), though they also eat considerable amounts of carrion. Komodo dragons also occasionally attack humans. Mating begins between May and August, and the eggs a ...
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Saniwa
''Saniwa'' is an extinct genus of varanid lizard that lived during the Eocene epoch. It is known from well-preserved fossils found in the Bridger and Green River Formations of Wyoming, United States. The type species ''S. ensidens'' was described in 1870 as the first fossil lizard known from North America. A second species, ''S.'' ''orsmaelensis'', is recognised from remains found in Europe. It is a close relative of '' Varanus'', the genus that includes monitor lizards. Description ''Saniwa'' measured . Like other varanid lizards, ''Saniwa'' had a long, pointed snout and nostrils placed farther back in the skull than most lizards and a tail that was almost twice as long as the body. Although similar in appearance to extant monitor lizards, ''Saniwa'' had many primitive traits, including teeth on its palate, a jugal bone beneath the eye that extended farther forward, and a suture between the frontal and parietal bones that was straight rather than curved. A study in 2018 by ...
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Anguimorpha
The Anguimorpha is a suborder of Squamata, squamates. The group was named by Fürbringer in 1900 to include all autarchoglossans closer to ''Varanus'' and ''Anguis'' than ''Scincus''. These lizards, along with iguanians and snakes, constitute the proposed "venom clade" Toxicofera of all venomous reptiles. Evolution The oldest widely accepted member of Anguimorpha is ''Dorsetisaurus'' from the Late Jurassic of Europe and North America. In 2022, the genus ''Cryptovaranoides'' was described from the latest Triassic (Rhaetian) of England. ''Cryptovaranoides'' was recovered in the study as a crown-group anguimorph, and therefore the oldest crown group-squamate, 35 million years older than any previously known crown-group squamate. However, a 2023 study found that ''Cryptovaranoides'' most likely represents an Archosauromorpha, archosauromorph that was only distantly related to squamates. Families Anguidae There are 9 genera found within the Anguidae family. They are characteriz ...
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Lanthanotus
The earless monitor lizard (''Lanthanotus borneensis'') is a semiaquatic, brown lizard native to the Southeast Asian island of Borneo. It is the monotypic, only living species in the family Lanthanotidae and it is related to the true monitor lizards. Taxonomy The earless monitor lizard was species description, described in 1878 by Franz Steindachner. The genus name ''Lanthanotus'' means "hidden ear" and the species name ''borneensis'' refers to its home island of Borneo. The uniqueness of the species was immediately recognized and Steindachner placed it in its own family, Lanthanotidae. In 1899, George Albert Boulenger relegated it to the family Helodermatidae, together with the Heloderma, beaded lizards and Gila monster, on the basis of morphological similarities. Further studies were conducted in the 1950s where it was found that although it is related to Helodermatidae, this relationship is relatively distant. The similarity is in part the result of convergent evolution and the ...
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Telmasaurus
''Telmasaurus'' is an extinct genus of varanoid lizard from the Late Cretaceous of Mongolia. Fossils have been found from the Djadokha and Barun Goyot Formations that date between the early and middle Campanian stage from approximately 80 to 71 million years ago. The type species ''Telmasaurus grangeri'' was named in 1943. Description ''Telmasaurus grangeri'' was named in 1943 and is now known from several fossils. Only two skulls are known, and the snout region is missing from both. Based on the size of these skulls, ''Telmasaurus'' was larger than other Cretaceous varanoid lizards, approaching the size of modern monitor lizards. Its skull is flatter than most other varanoids. The fusion of two bones in the skull called frontals links ''Telmasaurus'' with other early varanids like '' Paravaranus''. The lacrimal bone of ''Telmasaurus'' has a single hole in it, as do those of other early varanoids. The vertebrae of ''Telmasaurus'' are also known, and the way they attach to ...
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Earless Monitor Lizard
The earless monitor lizard (''Lanthanotus borneensis'') is a semiaquatic, brown lizard native to the Southeast Asian island of Borneo. It is the only living species in the family Lanthanotidae and it is related to the true monitor lizards. Taxonomy The earless monitor lizard was described in 1878 by Franz Steindachner. The genus name ''Lanthanotus'' means "hidden ear" and the species name ''borneensis'' refers to its home island of Borneo. The uniqueness of the species was immediately recognized and Steindachner placed it in its own family, Lanthanotidae. In 1899, George Albert Boulenger relegated it to the family Helodermatidae, together with the beaded lizards and Gila monster, on the basis of morphological similarities. Further studies were conducted in the 1950s where it was found that although it is related to Helodermatidae, this relationship is relatively distant. The similarity is in part the result of convergent evolution and they should be recognized as separate famil ...
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Ovoo Gurvel
''Ovoo gurvel'' is an extinct varanid lizard from the Late Cretaceous of Mongolia. It is one of the smallest and earliest monitor lizards. It was described in 2008. ''Ovoo'' possesses a pair of small bones in its skull that are not seen in any other lizard. Description and history ''Ovoo'' is only known from a fossilized skull cataloged as IGM 3/767 and designated the holotype. The skull was discovered in 2001 near the rich Ukhaa Tolgod fossil site in a locality known as Little Ukhaa. The deposits at Little Uhhaa date back to the Campanian stage of the Late Cretaceous. The generic name is derived from the name of a type of cairn called ovoos that are found along roads near Little Ukhaa. The specific name, ''gurvel'', comes from the Mongolian word for lizard. ''Ovoo'' was very small compared to living monitor lizards, with the exception of the Short-tailed monitor. Nevertheless, the structures of its skull are very similar to those of living monitors. ''Ovoo'' shares many simil ...
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Varanoidea
Varanoidea is a superfamily of lizards, including the well-known family Varanidae (the monitors and goannas). Also included in the Varanoidea are the Lanthanotidae ( earless monitor lizards), and the extinct Palaeovaranidae. Throughout their long evolutionary history, varanoids have exhibited great diversity, both in habitat and form. This superfamily includes the largest-known terrestrial lizard, Megalania (~5 meters), and the largest extant lizard, the Komodo dragon (''Varanus komodoensis'', ~3 meters). Evolution Either synonymous with, or a subgroup of, the group Platynota, the varanoids first appear in the fossil record in the latter part of Early Cretaceous, but possible varanoid ancestors have been traced back as far as Early Jurassic times. Among the earliest known varanoids are the monitor-like necrosaurids '' Palaeosaniwa canadensis'' from the Campanian (roughly 71–82 mya) of North America and '' Estesia mongoliensis'' and '' Telmasaurus grangeri'', both from the ...
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Savannah Monitor
The savannah monitor (''Varanus exanthematicus'') is a medium-sized species of monitor lizard native to Africa. The species is known as Bosc's monitor in Europe, since French scientist Louis Bosc first described the species. It belongs to the subgenus '' Polydaedalus''. Etymology The specific name ''exanthematicus'' is derived from the Greek word ''exanthem'' , meaning an eruption or blister of the skin. French botanist and zoologist Louis Augustin Guillaume Bosc originally described this lizard as ''Lacerta exanthematica'' in reference to the large oval scales on the back of its neck. The species was formerly known as ''Lacerta exanthematicus''. Description Savannah monitors are stoutly built, with relatively short limbs and toes, and skulls and dentition adapted to feed on hard-shelled prey. They are robust creatures, with powerful limbs for digging, powerful jaws and blunt, peglike teeth. Maximum size is rarely more than 100 cm. The skin coloration pattern varies accor ...
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Aiolosaurus
''Aiolosaurus'' is an extinct genus of monitor lizard from the Late Cretaceous of Mongolia. The type and only species, ''Aiolosaurus oriens'', was named in 2000 from Ukhaa Tolgod, a rich fossil site in the Campanian-age Djadochta Formation. Description and history ''Aiolosaurus'' was named in 2000 on the basis of a single holotype specimen cataloged as IGM 3/171. This specimen includes a partial skull and parts of the postcranial skeleton. ''Aiolosaurus'' is named after Aeolus, the Greek god of wind, while the specific name of ''A. oriens'' means "east." Diagnostic features of ''Aiolosaurus'' are found mainly in the skull. They include: *The division of the nasals into two bones (they form one bone in living monitors). *A small hole in the snout between the premaxilla and maxilla bones called the premaxillary fenestra. *The separation of the premaxilla and septomaxilla bones by a projection of the maxilla bone. *The small size of another hole in the snout called the sept ...
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Lizard
Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic islands, oceanic Archipelago, island chains. The grouping is Paraphyly, paraphyletic as some lizards are more closely related to snakes than they are to other lizards. Lizards range in size from chameleons and geckos a few centimeters long to the 3-meter-long Komodo dragon. Most lizards are quadrupedal, running with a strong side-to-side motion. Some lineages (known as "legless lizards") have secondarily lost their legs, and have long snake-like bodies. Some lizards, such as the forest-dwelling ''Draco (genus), Draco'', are able to glide. They are often Territory (animal), territorial, the males fighting off other males and signalling, often with bright colours, to attract mates and to intimidate rivals. Lizards are mainly carnivorous, often b ...
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Archaeovaranus
''Archaeovaranus'' (meaning "ancient '' Varanus''") is genus of varanid lizard from the early Eocene (Ypresian) Yuhuangding Formation of Hubei Province, China. The genus contains a single species, ''Archaeovaranus lii'', known from a nearly complete skeleton. The holotype, which includes an intact skull, is associated but disarticulated. ''Archaeovaranus'' fills a gap in the varanid fossil record, as it represents a stem-varanid from the early Eocene of East Asia, and is the closest known relative of '' Varanus''. Discovery and naming The holotype specimen of ''Archaeovaranus'', IVPP V 22770, was discovered at the Dajian locality of the Yuhuangding Formation near Danjiangkou, Hubei Province, China. The generic name, "''Archaeovaranus''," combines the Greek "archaīos," meaning "ancient," with a reference to the closely related ''Varanus''. "''Varanus''" is derived from the Arabic "waral," meaning "lizard beast." The specific name, "''lii''," honors the paleontologist Chuan ...
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