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Pteranodontids
The Pteranodontidae are a family of large pterosaurs from the Late Cretaceous of North America and possibly other continents including Europe and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head (most famously seen in ''Pteranodon'' itself). The spectacularly-crested ''Nyctosaurus'' is sometimes included in this family, though usually placed in its own family, the Nyctosauridae (Nicholson & Lydekker, 1889). Modern researchers differ in their use of the concept. S. Christopher Bennett and Alexander Kellner have concluded that ''Nyctosaurus'' was not a pteranodontid. In 1994 Bennett defined a clade Pteranodontidae, also including species of the Ornithocheiromorpha. However, this definition has not been accepted by other workers. Alexander Kellner, for example, named several additional species for specimens previously classified as ''Pteranodon'', and placed ''P. sternbergi'' in a distinc ...
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Ornithocheiromorpha
Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Ornithocheiromorphs have been discovered worldwide except Antarctica, though most genera have been recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by pteranodontians and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing ''Ornithocheirus'', but not ''Pteranodon''. Ornithocheiromorphs are considered to be some of the largest animals to have ever flown. Members of this group are also regarded to have some of the largest pterosaur wingspans, such as the one estimated for ...
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Nyctosaurus
''Nyctosaurus'' (meaning "night lizard") is a genus of nyctosaurid pterosaur from the Late Cretaceous period of what is now the Niobrara Formation of the mid-western United States, which, during the time ''Nyctosaurus'' was alive, was covered in an extensive shallow sea. Some remains belonging to a possible ''Nyctosaurus'' species called ''"N." lamegoi'' have been found in Brazil, though it likely belongs to a different genus '' Simurghia''. The genus ''Nyctosaurus'' has had numerous species referred to it, though how many of these may actually be valid requires further study. At least one species possessed an extraordinarily large antler-like cranial crest. ''Nyctosaurus'' was a mid-sized pterosaur that lived along the shores of the Niobrara Formation of the United States, which back then was within a large inland sea called the Western Interior Seaway. It has been suggested that it would have flown similar to modern-day soaring birds such as albatrosses, which consisted of fl ...
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Dawndraco
''Dawndraco'' is a controversial genus of pteranodontidae, pteranodontid pterosaur from the Late Cretaceous of North America. It is likely synonymous with the contemporary pteranodontid ''Geosternbergia.'' ''Dawndraco'' was named by Alexander Kellner, Alexander W.A. Kellner in 2010 in paleontology, 2010. The type species is ''Dawndraco kanzai''. The name of a biological genus, generic name combines the Dawn deity of the Iroquois with a Latin ''draco'', "dragon". The specific name (zoology), specific name refers to the Kaw people, Kanza tribe of Kansas. ''Dawndraco'' is based on the holotype specimen UALVP 24238, a partial skeleton including an almost complete skull and lower jaws. It was recovered in 1974 by Richard C. Fox and Allen Lindoe from rocks of the lower part of the Smoky Hill Chalk Member of the Niobrara Formation in Utica, Kansas. These rocks date to the late Coniacian or early Santonian stage, about 86 million years ago. The specimen was earlier referred to ''Ptera ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Aus ...
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Geosternbergia Sternbergi
''Pteranodon sternbergi'' is an extinct species of the pteranodontid pterodactyloid pterosaur genus ''Pteranodon'' from the Late Cretaceous geological period of North America. ''P. sternbergi'' was among the largest pterosaurs, with a wingspan of up to in males. It has been argued that ''P. sternbergi'' should be classified in a separate genus, ''Geosternbergia'', but this has not been followed by most other researchers. Discovery and history The first fossils of ''Pteranodon sternbergi'' were collected by American paleontologist George F. Sternberg in 1952 from the lower portion of the Niobrara Formation. The fossils of the animal looked similar to those of the species ''Pteranodon longiceps'', but the crests were set upright and in a slightly different position. In 1958, Sternberg and paleontologist Myrl V. Walker published a study about this peculiar find. In 1966, American paleontologist John Christian Harksen assigned the specimens found as a new species of ''Pteranod ...
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Pterosaur
Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or ' rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectivores or predators of small vertebrates. Later pte ...
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Volgadraco
''Volgadraco'' ("Volga River dragon") is a genus of pteranodontian pterosaur from the Upper Cretaceous of European Russia. Discovery and naming ''Volgadraco'' is known from lower beak (holotype SGU, no. 46/104a) and postcranial fragments from the early Campanian-age Rybushka Formation of Saratov, Russia. The size of this animal, and the development of blood supply in the lower jaw are intermediate between older Santonian or Turonian azhdarchids like ''Azhdarcho'' and '' Bakonydraco'' and later Maastrichtian azhdarchids like ''Quetzalcoatlus''. ''Volgadraco'' was described in 2008 by Averianov, Arkhangelsky, and Pervushov. The type species is ''V. bogolubovi'', the specific name honouring Russian paleontologist Nikolai Nikolaevich Bogolubov. Classification Averianov and colleagues initially assigned ''Volgadraco'' to the family Azhdarchidae, and considered the earlier named genus '' Bogolubovia'' to be a ''nomen dubium'' that is potentially identical to ''Volgadraco''. In contras ...
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Tethydraco
''Tethydraco'' is a genus of pterodactyloid pterosaur that lived during the Late Cretaceous period (Maastrichtian stage) of what is now the area of present Morocco, about 66 million years ago. ''Tethydraco'' was originally assigned to the family Pteranodontidae. Some researchers argued that subsequently described material suggests that it may have been an azhdarchid, and possibly synonymous with '' Phosphatodraco'', though this has been disputed. The type and only species is ''T. regalis''. Discovery and naming Since 2015, a group of paleontologists has been acquiring pterosaur fossils from commercial Moroccan fossil traders, who obtain these from workers in the phosphate mines on the Khouribga plateau, which is located within the Ouled Abdoun Basin. The purpose of this project is to determine pterosaur diversity in the latest Cretaceous. From this stage, no Konservat-Lagerstätten are known, sites combining a large variety of species with exceptional preservation. It is in ...
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Pteranodon
''Pteranodon'' (; from and ) is a genus of pterosaur that included some of the largest known flying reptiles, with ''P. longiceps'' having a wingspan of over . They lived during the late Cretaceous geological period of North America in present-day Kansas, Nebraska, Wyoming, South Dakota and Alabama. More fossil specimens of ''Pteranodon'' have been found than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved with nearly complete skulls and articulated skeletons. It was an important part of the animal community in the Western Interior Seaway. When the first fossils of ''Pteranodon'' were found, they were assigned to toothed pterosaur genera, ''Ornithocheirus'' and ''Pterodactylus''. In 1876, Othniel Charles Marsh recognised it as a genus of its own, making particular note of its complete lack of teeth, which at the time was unique among pterosaurs. Over the decades, multiple species would be assigned to ''Pteranodon'', though today, o ...
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Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (stratigraphy), Series, the Cretaceous geologic period, Period or system (stratigraphy), System, and of the Mesozoic geologic era, Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). It is named after the city of Maastricht, the capital and largest city of the Limburg (Netherlands), Limburg province in the Netherlands. The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary period, Tertiary extinction event) occurred at the end of this age. In this extinction event, mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-kn ...
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Geosternbergia
''Pteranodon sternbergi'' is an extinct species of the pteranodontid pterodactyloid pterosaur genus ''Pteranodon'' from the Late Cretaceous geological period of North America. ''P. sternbergi'' was among the largest pterosaurs, with a wingspan of up to in males. It has been argued that ''P. sternbergi'' should be classified in a separate genus, ''Geosternbergia'', but this has not been followed by most other researchers. Discovery and history The first fossils of ''Pteranodon sternbergi'' were collected by American paleontologist George F. Sternberg in 1952 from the lower portion of the Niobrara Formation. The fossils of the animal looked similar to those of the species ''Pteranodon longiceps'', but the crests were set upright and in a slightly different position. In 1958, Sternberg and paleontologist Myrl V. Walker published a study about this peculiar find. In 1966, American paleontologist John Christian Harksen assigned the specimens found as a new species of ''Pteranodon' ...
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Pteranodon Longiceps
''Pteranodon'' (; from and ) is a genus of pterosaur that included some of the largest known flying reptiles, with ''P. longiceps'' having a wingspan of over . They lived during the late Cretaceous geological period of North America in present-day Kansas, Nebraska, Wyoming, South Dakota and Alabama. More fossil specimens of ''Pteranodon'' have been found than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved with nearly complete skulls and articulated skeletons. It was an important part of the animal community in the Western Interior Seaway. When the first fossils of ''Pteranodon'' were found, they were assigned to toothed pterosaur genera, ''Ornithocheirus'' and ''Pterodactylus''. In 1876, Othniel Charles Marsh recognised it as a genus of its own, making particular note of its complete lack of teeth, which at the time was unique among pterosaurs. Over the decades, multiple species would be assigned to ''Pteranodon'', though today, o ...
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