Volgadraco
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''Volgadraco'' ("
Volga River The Volga (, ) is the longest river in Europe and the longest endorheic basin river in the world. Situated in Russia, it flows through Central Russia to Southern Russia and into the Caspian Sea. The Volga has a length of , and a catchment ...
dragon A dragon is a Magic (supernatural), magical legendary creature that appears in the folklore of multiple cultures worldwide. Beliefs about dragons vary considerably through regions, but European dragon, dragons in Western cultures since the Hi ...
") is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
pteranodontia Pteranodontia is an extinct group of ornithocheiroid pterodactyloid pterosaurs. It lived during the Late Cretaceous (Turonian to Maastrichtian stages) of North America, South America, Europe and Africa. They were some of the most advanced pterosa ...
n
pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
from the
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cret ...
of European
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
.


Discovery and naming

''Volgadraco'' is known from lower beak (
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
SGU, no. 46/104a) and
postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...
l fragments from the early
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
-age Rybushka Formation of
Saratov Saratov ( , ; , ) is the largest types of inhabited localities in Russia, city and administrative center of Saratov Oblast, Russia, and a major port on the Volga River. Saratov had a population of 901,361, making it the List of cities and tow ...
, Russia. The size of this animal, and the development of blood supply in the lower jaw are intermediate between older
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...
or
Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...
azhdarchids like ''
Azhdarcho ''Azhdarcho'' is a genus of azhdarchid pterosaur from the late Cretaceous Period (geology), Period of the Bissekty Formation (middle Turonian stage, about 92 million years ago) of Uzbekistan, as well as the Zhirkindek Formation of Kazakhstan a ...
'' and '' Bakonydraco'' and later
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
azhdarchids like ''
Quetzalcoatlus ''Quetzalcoatlus'' () is a genus of azhdarchid pterosaur that lived during the Maastrichtian age of the Late Cretaceous in North America. The Type (biology), type specimen, recovered in 1971 from the Javelina Formation of Texas, United States, ...
''. ''Volgadraco'' was described in 2008 by Averianov, Arkhangelsky, and Pervushov. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''V. bogolubovi'', the
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honouring Russian paleontologist Nikolai Nikolaevich Bogolubov.


Classification

Averianov and colleagues initially assigned ''Volgadraco'' to the family
Azhdarchidae Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cre ...
, and considered the earlier named genus '' Bogolubovia'' to be a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
'' that is potentially identical to ''Volgadraco''. In contrast, Longrich and colleagues in 2018 had recovered it within the family
Nyctosauridae Nyctosauridae (meaning "night lizards" or "bat lizards") is a family of specialized soaring pterosaurs of the late Cretaceous Period of North America, Africa, and possibly other continents including South America. It was named in 1889 by Henry ...
, the cladogram of which can be seen below: In 2020, Averianov and Arkhangelsky reconsidered both ''Bogolubovia'' and ''Volgadraco'' as members of the family
Pteranodontidae The Pteranodontidae are a Family (biology), family of large pterosaurs from the Late Cretaceous of North America and possibly other continents including Europe and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a ...
. In 2022, Averianov and Kurin considered both ''Volgadraco'' and ''Bogolubovia'' as valid pteranodontids based on a new specimen of ''Bogolubovia''. In the same year, however, ''Volgadraco'' was considered to be a pteranodontian both outside the
Nyctosauridae Nyctosauridae (meaning "night lizards" or "bat lizards") is a family of specialized soaring pterosaurs of the late Cretaceous Period of North America, Africa, and possibly other continents including South America. It was named in 1889 by Henry ...
and
Pteranodontidae The Pteranodontidae are a Family (biology), family of large pterosaurs from the Late Cretaceous of North America and possibly other continents including Europe and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a ...
based on phylogenetic analysis, the cladogram of which can be seen below:


See also

*
Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and Biological taxonomy, taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic Era (ge ...
* List of pterosaurs


References

{{Portal bar, Paleontology, Russia Pteranodontia Late Cretaceous pterosaurs of Europe Fossil taxa described in 2008 Campanian genera Cretaceous Russia Fossils of Russia