Protorothyridid
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Protorothyridid
Protorothyrididae is an extinct family of small, lizard-like reptiles belonging to Eureptilia. Their skulls did not have fenestrae, like the more derived diapsids. Protorothyridids lived from the Late Carboniferous to Early Permian periods, in what is now North America. Many genera of primitive reptiles were thought to be protorothyridids. ''Brouffia'', ''Coelostegus'', ''Paleothyris'' and ''Hylonomus'', for example, were found to be more basal eureptiles in Muller and Reisz (2006), making the family as historically defined paraphyletic, though three genera, ''Protorothyris, Anthracodromeus,'' and ''Cephalerpeton'', were recovered as a monophyletic group. ''Anthracodromeus'', ''Paleothyris'', and ''Protorothyris'' were recovered as a monophyletic group in Ford and Benson (2020) (who did not sample ''Cephalerpeton''), who recovered them as more derived than captorhinids and ''Hylonomus'', but less so than araeoscelidians. ''Anthracodromeus'' is the earliest known reptile to displa ...
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Protorothyrididae
Protorothyrididae is an extinct family (biology), family of small, lizard-like reptiles belonging to Eureptilia. Their skulls did not have Fenestra (anatomy), fenestrae, like the more derived diapsids. Protorothyridids lived from the Late Carboniferous to Cisuralian, Early Permian periods, in what is now North America. Many genera of primitive reptiles were thought to be protorothyridids. ''Brouffia'', ''Coelostegus'', ''Paleothyris'' and ''Hylonomus'', for example, were found to be more basal (phylogenetics), basal eureptiles in Muller and Reisz (2006), making the family as historically defined paraphyletic, though three genera, ''Protorothyris, Anthracodromeus,'' and ''Cephalerpeton'', were recovered as a monophyletic group. ''Anthracodromeus'', ''Paleothyris'', and ''Protorothyris'' were recovered as a monophyletic group in Ford and Benson (2020) (who did not sample ''Cephalerpeton''), who recovered them as more derived than captorhinids and ''Hylonomus'', but less so than arae ...
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Protorothyris
''Protorothyris'' is an extinct genus of Early Permian protorothyridid known from Texas and West Virginia of the United States. It was first named by Llewellyn Ivor Price in 1937 and the type species is ''Protorothyris archeri''. ''P. archeri'' is known from the holotype MCZ 1532, a three-dimensionally preserved skull and from the referred specimens, which come from four additional individuals, MCZ 2147-2150. All specimens were collected in the Cottonwood Creek site, from the Archer City Formation, Texas, dating to the Asselian stage of the Cisuralian epoch, about 299–294.6 million years ago. A second species, ''P. morani'', was first named by Alfred Sherwood Romer in 1952 with its own generic name, ''Melanothyris''. In 1973, J. Clark and Robert L. Carroll recombined ''P. morani'' as a ''Protorothyris'' species. It is known from the holotype CM 8617, a three-dimensionally preserved skull. It was collected in the Blacksville site, from the Washington Formation ...
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Anthracodromeus
''Anthracodromeus'' is an extinct genus of Late Carboniferous (late Westphalian (stage), Westphalian stage) "Protorothyrididae, protorothyridid" tetrapods known from Ohio. It is known from the holotype American Museum of Natural History, AMNH 6940, a nearly complete skeleton. It was collected in the Linton site in Jefferson County, Ohio, Jefferson County, from the Allegheny Group. ''A. longipes'' was first assigned by Edward Drinker Cope in 1875 in paleontology, 1875 to a species of ''Sauropleura''. The genus was first named by Robert L. Carroll and Donald Baird in 1972 in paleontology, 1972 and the type species is ''Anthracodromeus longipes''. It is amongst the oldest known tetrapods to display adaptations for climbing. Features and environment ''Anthracodromeus'' was between 15 and 20 cm long when alive, with about 5 cm of this being tail. It had a long body, with 27 vertebrae between the five cervical vertebrae and the sacrum. Each vertebra had a short pair of ...
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Cephalerpeton
''Cephalerpeton'' is an extinct genus of " protorothyridid" tetrapods known from the Late Carboniferous (late Westphalian stage) of Illinois. It is known from the holotype YPM 796, a partial skeleton. It was collected in the Mazon Creek site, from the Francis Creek Shale Member of the Carbondale Formation. It was first named by R. L. Moodie in 1912 as an amphibamid amphibian and the type species is ''Cephalerpeton ventriarmatum''. It was first assigned to Protorothyrididae by Robert L. Carroll Robert "Bob" Lynn Carroll (May 5, 1938 – April 7, 2020) was an American–Canadian vertebrate paleontologist who specialised in Paleozoic and Mesozoic amphibians and reptiles. Biography Carroll was an only child and grew up on a farm nea ... and Donald Baird in 1972 and this placement has been widely accepted. References Protorothyrididae Pennsylvanian tetrapods of North America Paleontology in Illinois Fossil taxa described in 1912 {{paleo-reptile-stub ...
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Captorhinids
Captorhinidae is an extinct family of tetrapods, traditionally considered primitive reptiles, known from the late Carboniferous to the Late Permian. They had a cosmopolitan distribution across Pangea. Description Captorhinids are a clade of small to very large lizard-like animals that date from the Late Carboniferous through the Permian. Their skulls were much stronger than those of their relatives, the protorothyridids, and had teeth that were better able to deal with tough plant material. The postcranial skeleton is similar to those of seymouriamorphs and diadectomorphs; these animals were grouped together with the captorhinids in the order Cotylosauria as the first reptiles in the early 20th century, but are now usually regarded as stem-amniotes no closer to reptiles than to mammals. Captorhinids have broad, robust skulls that are generally triangular in shape when seen in dorsal view. The premaxillae are characteristically downturned. The largest captorhinid, the herbiv ...
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Pennsylvanian (geology)
The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, on the International Commission on Stratigraphy, ICS geologic timescale, the younger of two period (geology), subperiods of the Carboniferous Period (or the upper of two system (stratigraphy), subsystems of the Carboniferous System). It lasted from roughly . As with most other geochronology, geochronologic units, the stratum, rock beds that define the Pennsylvanian are well identified, but the exact date of the start and end are uncertain by a few hundred thousand years. The Pennsylvanian is named after the U.S. state of Pennsylvania, where the coal Bed (geology), beds of this age are widespread. The division between Pennsylvanian and Mississippian (geology), Mississippian comes from North American stratigraphy. In North America, where the early Carboniferous beds are primarily marine limestones, the Pennsylvanian was in the past treated as a full-fledged geologic period between the Mississippian ...
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Paraphyletic
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor exc ...
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Monophyletic
In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent common ancestor (or more precisely an ancestral population), i.e. excludes non-descendants of that common ancestor # the grouping contains all the descendants of that common ancestor, without exception Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic'' grouping meets 1. but not 2., thus consisting of the descendants of a common ancestor, excepting one or more monophyletic subgroups. A '' polyphyletic'' grouping meets neither criterion, and instead serves to characterize convergent relationships of biological features rather than genetic relationships – for example, night-active primates, fruit trees, or aquatic insects. As such, these characteristic features of a polyphyletic grouping ...
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Araeoscelidians
Araeoscelidia or Araeoscelida is a clade of extinct tetrapods (traditionally classified as diapsid reptiles) superficially resembling lizards, extending from the Late Carboniferous to the Early Permian. The group contains the genera ''Araeoscelis'', ''Petrolacosaurus'', the possibly aquatic '' Spinoaequalis'', and less well-known genera such as ''Kadaliosaurus'' and ''Zarcasaurus''. This clade is usually considered to be the sister group to all (currently known) later diapsids. Description Araeoscelidians were small animals (less than one meter in length) looking somewhat like lizards, though they are only distantly related to true lizards. They differ from other, earlier sauropsids by their slender limbs, their elongated tail, and of course by the presence of two temporal openings, the feature defining the diapsid condition. In ''Araeoscelis'', only the upper temporal opening remains, thus resulting in a derived euryapsid condition. Genera Araeoscelidia includes well-known gen ...
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Crown Group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crow ...
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Arboreal Locomotion
Arboreal locomotion is the animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolution, evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The habitats pose numerous mechanical challenges to animals moving through them and lead to a variety of anatomical, behavioral and ecological consequences as well as variations throughout different species.Matt Cartmill, Cartmill, M. (1985). "Climbing". pp. 73–88 ''In'': Hildebrand, Milton; Bramble, Dennis M.; species:Karel Frederik Liem, Liem, Karel F.; David B. Wake, Wake, David B. (editors) (1985). ''Functional Vertebrate Morphology''. Cambridge, Massachusetts: Harvard University Press, Belknap Press. 544 pp. . Furthermore, many of these same principles may be applied to climbing without trees, such as on rock piles or mountains. Some animals are exclusively arboreal in habitat, such as tree snails. Biomechanic ...
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