Phragmocone
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Phragmocone
The phragmocone is the chambered portion of the shell of a cephalopod. It is divided by septa into camerae. In most nautiloids and ammonoids, the phragmocone is a long, straight, curved, or coiled structure, in which the camerae are linked by a siphuncle which determines buoyancy by means of gas exchange. Despite this benefit, such a large shell adds to the mass of the animal, and hence is disadvantageous in catching fast-moving prey. Some nautiloids, such as the Silurian Ascocerida, dropped the phragmocone upon maturity, presumably to increase speed and maneuverability. They thus became the early Paleozoic equivalent of coleoids. The early coleoids and belemnoids adopted a different approach: the phragmocone was retained but became internal and reduced. In general the shell in cephalopods tends to be vestigial or absent. Fossil record Being the only biomineralised part of most cephalopods, the phragmocone is typically the only part to enter the fossil record. It is s ...
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Belemnoidea
Belemnoids are an extinct group of marine cephalopod, very similar in many ways to the modern squid. Like them, the belemnoids possessed an ink sac, but, unlike the squid, they possessed ten Cephalopod arm, arms of roughly equal length, and no tentacles. The name "belemnoid" comes from the Greek word βέλεμνον, ''belemnon'' meaning "a dart or arrow" and the Greek word είδος, ''eidos'' meaning "form". Belemnoids include Belemnitida, belemnites (which belong to the Order (biology), order Belemnitida), aulacocerids (order Aulacocerida), phragmoteuthids (order Phragmoteuthida), and diplobelids (order Diplobelida). Belemnoidea has been suggested to be paraphyletic by some authors. Occurrence Belemnoids were numerous during the Jurassic and Cretaceous periods, and their fossils are abundant in Mesozoic marine rocks, often accompanying their cousins the ammonites. The belemnoids become extinct at the end of the Cretaceous period along with the ammonites. The belemnoids' or ...
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Ammonoidea
Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family Nautilidae). The earliest ammonoids appeared during the Emsian stage of the Early Devonian (410.62 million years ago), with the last species vanishing during or soon after the Cretaceous–Paleogene extinction event (66 million years ago). They are often called ammonites, which is most frequently used for members of the order Ammonitida, the only remaining group of ammonoids from the Jurassic up until their extinction. Ammonoids exhibited considerable diversity over their evolutionary history, with over 10,000 species having been described. Ammonoids are excellent index fossils, and they have been frequently used to link rock layers in which a particular species or genus is found to specific Geologic time scale, geologic time periods. Their ...
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Nautiloid
Nautiloids are a group of cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and species rich, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). No orthoconic and only a handful of coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-desce ...
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Septa (biology)
Septa (singular septum) are thin walls or partitions between the internal chambers (camera (cephalopod), camerae) of the Exoskeleton, shell of a cephalopod, namely nautiloids or Ammonoidea, ammonoids. As the creature grows, its body moves forward in the shell to a new living chamber, secreting septa behind it. This adds new chambers to the shell, which can be clearly seen in cross-sections of the shell of the living nautilus, or in ammonoid and nautiloid fossils. The septa are attached to the inside wall of the shell, thus dividing the phragmocone into camerae. Where the septum meets the shell a suture (anatomical), suture line forms; in some ammonoids these lines became extremely complex and elaborate, providing strength without the necessity of added weight. Elaborate sutures allowed for thinner shells, and hence less time needed for shell growth and less time spent in the vulnerable juvenile stage. The nature and structure of the septa, as with the camerae, and siphuncle, ...
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Camera (cephalopod)
Camerae (singular camera) are the spaces or chambers enclosed between two adjacent Septa (biology), septa in the phragmocone of a nautiloid or Ammonoidea, ammonoid cephalopod, cephalopod molluscus. These can be seen in cross-sections of a nautilus shell and in the polished cross-sections of ammonites. In life these chambers are filled with gas, mediated by the siphuncle, and used to control buoyancy. Some Palaeozoic nautiloid genera, especially those with Orthocone, long, straight shells, are distinguished by cameral deposits. These were accumulations of calcium carbonate secreted in the empty chambers of the shell, used for Ballast tanks, ballast and control of buoyancy. The nature and form of these deposits are very useful in nautiloid classification. References

* {{DEFAULTSORT:Camera (Anatomy) Cephalopod zootomy ...
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Exoskeleton
An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the internal organs, in contrast to an internal endoskeleton (e.g. human skeleton, that of a human) which is enclosed underneath other soft tissues. Some large, hard and non-flexible protective exoskeletons are known as mollusc shell, shell or armour (anatomy), armour. Examples of exoskeletons in animals include the arthropod exoskeleton, cuticle skeletons shared by arthropods (insects, chelicerates, myriapods and crustaceans) and tardigrades, as well as the corallite, skeletal cups formed by hardened secretion of stony corals, the test (biology), test/tunic of sea squirts and sea urchins, and the prominent mollusc shell shared by snails, bivalvia, clams, tusk shells, chitons and nautilus. Some vertebrate animals, such as the turtle, have both an endoskeleton and a turtle shell, protective exoskeleton. Role Exoskeletons c ...
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Cephalopod
A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symmetry, bilateral body symmetry, a prominent head, and a set of cephalopod arm, arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt Cephalopod ink, ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, Extant taxon, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus (genus), Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, where ...
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Siphuncle
The siphuncle is a strand of biological tissue, tissue passing longitudinally through the mollusc shell, shell of a cephalopod mollusc. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ''Spirula''. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the Septum (cephalopod), septa (walls) dividing the Camera (cephalopod), camerae (chambers). Some older studies have used the term siphon for the siphuncle, though this naming convention is uncommon in modern studies to prevent confusion with a Siphon (mollusc), mollusc organ of the same name. Function The siphuncle is used primarily in emptying water from new chambers as the shell grows. To perform this task, the cephalopod increases the saltiness of the blood in the siphuncle, ...
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Buoyancy
Buoyancy (), or upthrust, is the force exerted by a fluid opposing the weight of a partially or fully immersed object (which may be also be a parcel of fluid). In a column of fluid, pressure increases with depth as a result of the weight of the overlying fluid. Thus, the pressure at the bottom of a column of fluid is greater than at the top of the column. Similarly, the pressure at the bottom of an object submerged in a fluid is greater than at the top of the object. The pressure difference results in a net upward force on the object. The magnitude of the force is proportional to the pressure difference, and (as explained by Archimedes' principle) is equivalent to the weight of the fluid that would otherwise occupy the submerged volume of the object, i.e. the Displacement (fluid), displaced fluid. For this reason, an object with average density greater than the surrounding fluid tends to sink because its weight is greater than the weight of the fluid it displaces. If the objec ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods, arachnids and hexapods) ...
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Ascocerida
The Ascocerida are comparatively small, bizarre Orthoceratoids known only from Ordovician and Silurian sediments in Europe and North America, uniquely characterized by a deciduous conch consisting of a longer juvenile portion and an inflated short adult portion that separate sometime in maturity.W.M . Furnish and Brian F Glenister; Nautiloidea-Ascocerida, K261-K277; Treatise on Invertebrate Paleontology, part K, Nautiloidea; Geological Society of America and the University of Kansas Press, 1964 Morphology The juvenile portion of an ascocerid consists typically of a narrow, circular cyrtocone that underwent periodic truncation. Cumulative length, including broken off segments, may have reached about in the largest individuals. The siphuncle is located halfway between the shell axis and the venter, is thin walled and tubular with short, orthochoanitic septal necks and segments that are only slightly inflated. The juvenile portion is known as the deciduous conch, as it is shed in a ...
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