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Hybodonts
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like cartilaginous fish (chondrichthyans) which existed from the late Devonian to the Late Cretaceous. Hybodonts share a close common ancestry with modern sharks and Batoidea, rays (Neoselachii) as part of the clade Euselachii. They are distinguished from other chondrichthyans by their distinctive fin spines and cephalic spines present on the heads of males. An ecologically diverse group, they were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct. Etymology The term hybodont comes from the Greek word ''ὕβος'' or ''ὑβός'' meaning hump or hump-backed and ''ὀδούς, ὀδοντ'' meaning tooth. This n ...
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Elasmobranchii
Elasmobranchii () is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks ( division Selachii), and batomorphs (division Batomorphi, including rays, skates, and sawfish). Members of this subclass are characterised by having five to seven pairs of gill slits opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil. The definition of the clade is unclear with respect to fossil chondrichthyans. Some authors consider it as equivalent to Neoselachii (the crown group clade including modern sharks, rays, and all o ...
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Holocephali
Holocephali (Sometimes spelled Holocephala; Romanization of Greek, Greek for "complete head" in reference to the fusion of Palatoquadrate, upper jaw with the rest of the skull) is a Subclass (biology), subclass of Chondrichthyes, cartilaginous fish. While the only living holocephalans are three families within a single Order (biology), order which together are commonly known as chimaeras, the group includes many extinct orders and was far more diverse during the Paleozoic and Mesozoic Era (geology), eras. The earliest known fossils of holocephalans date to the Middle Devonian period, and the group likely reached its peak diversity during the following Carboniferous period. Molecular clock studies suggest that the subclass diverged from its closest relatives, Elasmobranchii, elasmobranchs such as sharks and Batomorphi, rays, during the Early Devonian or Silurian period. Extinct holocephalans are typically divided into a number of orders, although the interrelationships of these gro ...
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Shark
Sharks are a group of elasmobranch cartilaginous fish characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the Division (taxonomy), division Selachii and are the sister group to the Batoidea, Batomorphi (Batoidea, rays and skate (fish), skates). Some sources extend the term "shark" as an informal category including Extinction, extinct members of Chondrichthyes (cartilaginous fish) with a shark-like morphology, such as hybodonts. Shark-like chondrichthyans such as ''Cladoselache'' and ''Doliodus'' first appeared in the Devonian Period (419–359 million years), though some fossilized chondrichthyan-like scales are as old as the Ordovician, Late Ordovician (458–444 million years ago). The earliest confirmed modern sharks (Selachii) are known from the Early Jurassic around , with the oldest known member being ''Agaleus'', though records of true shar ...
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Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. The Jurassic constitutes the second and middle period of the Mesozoic, Mesozoic Era as well as the eighth period of the Phanerozoic, Phanerozoic Eon and is named after the Jura Mountains, where limestone strata from the period were first identified. The start of the Jurassic was marked by the major Triassic–Jurassic extinction event, associated with the eruption of the Central Atlantic magmatic province, Central Atlantic Magmatic Province (CAMP). The beginning of the Toarcian Age started around 183 million years ago and is marked by the Toarcian Oceanic Anoxic Event, a global episode of Anoxic event, oceanic anoxia, ocean acidification, and elevated global temperatures associated with extinctions, likely caused by the eruption of the Kar ...
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Sharks
Sharks are a group of elasmobranch cartilaginous fish characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the Division (taxonomy), division Selachii and are the sister group to the Batoidea, Batomorphi (Batoidea, rays and skate (fish), skates). Some sources extend the term "shark" as an informal category including Extinction, extinct members of Chondrichthyes (cartilaginous fish) with a shark-like morphology, such as hybodonts. Shark-like chondrichthyans such as ''Cladoselache'' and ''Doliodus'' first appeared in the Devonian Period (419–359 million years), though some fossilized chondrichthyan-like scales are as old as the Ordovician, Late Ordovician (458–444 million years ago). The earliest confirmed modern sharks (Selachii) are known from the Early Jurassic around , with the oldest known member being ''Agaleus'', though records of true shar ...
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Asteracanthus
''Asteracanthus'' (from , 'star' and , 'spine') is an extinct genus of hybodont, known from the Middle Jurassic (Bathonian) to the Early Cretaceous (Valanginian). Description ''Asteracanthus'' was among the largest known hybodontiforms, reaching a length of . The dentition of ''Astercanthus'' is high crowned and multicusped. The low-rounded teeth previously attributed to the genus actually belong to '' Strophodus''. Like ''Strophodus'', the fin spines are covered in tubercles, rather than the ribbing typical of other hybodont fin spines. Fossil records The genus as currently circumscribed dates from the Middle Jurassic (Bathonian) to Early Cretaceous (Valanginian) of Europe, including Germany, Switzerland, England and France. A complete skeleton was described in 2021 from the Late Jurassic (Tithonian) aged Solnhofen Limestone. Previously considered synonymous, the genus '' Strophodus'' (Middle Triassic-Late Cretaceous) is now considered distinct, with the teeth of ''Asterac ...
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Dracopristis Hoffmanorum
''Dracopristis'' (meaning 'dragon shark') is an extinct genus of ctenacanth (a group of shark-like cartilaginous fish) that lived during the Carboniferous period in North America, around 307 million years ago. The species was discovered in the Kinney Brick Quarry in New Mexico, USA. Like many fossils from the site, the fossils of ''Dracopristis'' are very well-preserved. A single species is known, ''Dracopristis hoffmanorum'', which is named in honor of Ralph and Jeanette Hoffman, the owners of the quarry. Prior to being scientific named, ''D. hoffmanorum'' was informally referred to as the "Godzilla shark". ''Dracopristis'' possessed large dermal denticles along its head, along with rows of short, multi-cusped teeth in its jaws and very large spines on its dorsal fins, the latter of which inspired the name of the genus. The dorsal spines of the type specimen are about in length, while the entire body was approximately in length. Its large spines were likely used for def ...
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Ctenacanthiformes
Ctenacanthiformes is an extinct order of cartilaginous fish. They possessed ornamented fin spines at the front of their dorsal fins and cladodont-type dentition, that is typically of a grasping morphology, though some taxa developed cutting and gouging tooth morphologies. Some ctenacanths are thought to have reached sizes comparable to the great white shark, with body lengths of up to and weights of , while others reached lengths of only . The earliest ctenacanths appeared during the Frasnian stage of the Late Devonian (~383-372 million years ago), with the group reaching their greatest diversity during the Early Carboniferous ( Mississippian), and continued to exist into at least the Middle Permian (Guadalupian). Some authors have suggested members of the family Ctenacanthidae may have survived into the Cretaceous based on teeth found in deep water deposits of Valanginian age in France and Austria, however, other authors contend that the similarity of these teeth to Paleozoic ct ...
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Chimaeras
Chimaeras are Chondrichthyes, cartilaginous fish in the order (biology), order Chimaeriformes (), known informally as ghost sharks, rat fish (not to be confused with rattails), spookfish, or rabbit fish; the last two names are also applied to Barreleye, Opisthoproctidae and Rabbitfish, Siganidae, respectively. At one time a "diverse and abundant" group (based on the fossil record), their closest living relatives are sharks and ray (fish), rays, though their last common ancestor with them lived nearly 400 million years ago. Living species (aside from plough-nose chimaeras) are largely confined to deep water. Anatomy Chimaeras are soft-bodied, shark-like fish with bulky heads and long, tapered tails; measured from the tail, they can grow up to in length. Like other members of the class Chondrichthyes, chimaera skeletons are entirely cartilaginous, or composed of cartilage. Males use forehead denticles to grasp a female by a fin during copulation. The Branchial arch, gill arche ...
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