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Fractionation Of Carbon Isotopes In Oxygenic Photosynthesis
Photosynthesis converts carbon dioxide to carbohydrates via several metabolic pathways that provide energy to an organism and preferentially react with certain stable isotopes of carbon. The selective enrichment of one stable isotope over another creates distinct Isotope fractionation, isotopic fractionations that can be measured and correlated among oxygenic phototrophs. The degree of carbon isotope fractionation is influenced by several factors, including the metabolism, anatomy, growth rate, and environmental conditions of the organism. Understanding these variations in carbon fractionation across species is useful for Biogeochemistry, biogeochemical studies, including the reconstruction of paleoecology, plant evolution, and the characterization of food chains. Oxygenic photosynthesis is a metabolic pathway facilitated by autotrophs, including plants, algae, and cyanobacteria. This pathway converts inorganic carbon dioxide from the atmosphere or aquatic environment into carbohydr ...
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Photosynthesis
Photosynthesis ( ) is a system of biological processes by which photosynthetic organisms, such as most plants, algae, and cyanobacteria, convert light energy, typically from sunlight, into the chemical energy necessary to fuel their metabolism. ''Photosynthesis'' usually refers to oxygenic photosynthesis, a process that produces oxygen. Photosynthetic organisms store the chemical energy so produced within intracellular organic compounds (compounds containing carbon) like sugars, glycogen, cellulose and starches. To use this stored chemical energy, an organism's cells metabolize the organic compounds through cellular respiration. Photosynthesis plays a critical role in producing and maintaining the oxygen content of the Earth's atmosphere, and it supplies most of the biological energy necessary for complex life on Earth. Some bacteria also perform anoxygenic photosynthesis, which uses bacteriochlorophyll to split hydrogen sulfide as a reductant instead of water, p ...
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Rate Limiting Step
In chemical kinetics, the overall rate of a reaction is often approximately determined by the slowest step, known as the rate-determining step (RDS or RD-step or r/d step) or rate-limiting step. For a given reaction mechanism, the prediction of the corresponding rate equation (for comparison with the experimental rate law) is often simplified by using this approximation of the rate-determining step. In principle, the time evolution of the reactant and product concentrations can be determined from the set of simultaneous rate equations for the individual steps of the mechanism, one for each step. However, the analytical solution of these differential equations is not always easy, and in some cases numerical integration may even be required. The hypothesis of a single rate-determining step can greatly simplify the mathematics. In the simplest case the initial step is the slowest, and the overall rate is just the rate of the first step. Also, the rate equations for mechanisms with a ...
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NADP+
Nicotinamide adenine dinucleotide phosphate, abbreviated NADP or, in older notation, TPN (triphosphopyridine nucleotide), is a cofactor used in anabolic reactions, such as the Calvin cycle and lipid and nucleic acid syntheses, which require NADPH as a reducing agent ('hydrogen source'). NADPH is the reduced form, whereas NADP is the oxidized form. NADP is used by all forms of cellular life. NADP is essential for life because it is needed for cellular respiration. NADP differs from NAD by the presence of an additional phosphate group on the 2' position of the ribose ring that carries the adenine moiety. This extra phosphate is added by NAD+ kinase and removed by NADP+ phosphatase. Biosynthesis NADP In general, NADP+ is synthesized before NADPH is. Such a reaction usually starts with NAD+ from either the de-novo or the salvage pathway, with NAD+ kinase adding the extra phosphate group. ADP-ribosyl cyclase allows for synthesis from nicotinamide in the salvage pathway, a ...
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Carbon Fixation
Biological carbon fixation, or сarbon assimilation, is the Biological process, process by which living organisms convert Total inorganic carbon, inorganic carbon (particularly carbon dioxide, ) to Organic compound, organic compounds. These organic compounds are then used to store energy and as structures for other Biomolecule, biomolecules. Carbon is primarily fixed through photosynthesis, but some organisms use chemosynthesis in the absence of sunlight. Chemosynthesis is carbon fixation driven by chemical energy rather than from sunlight. The process of biological carbon fixation plays a crucial role in the global carbon cycle, as it serves as the primary mechanism for removing from the atmosphere and incorporating it into Biomass (ecology), living biomass. The primary production of organic compounds allows carbon to enter the biosphere. Carbon is considered essential for life as a base element for building organic compounds. The flow of carbon from the Earth's atmosphere, ...
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Belemnitida
Belemnitida (or belemnites) is an extinct order (biology), order of squid-like cephalopods that existed from the Late Triassic to Late Cretaceous (And possibly the Eocene). Unlike squid, belemnites had an internal skeleton that made up the cone. The parts are, from the arms-most to the tip, the tongue-shaped pro-ostracum, the conical phragmocone, and the pointy guard. The calcite, calcitic guard is the most common belemnite remain. Belemnites, in life, are thought to have had 10 hooked arms and a pair of fins on the guard. The chitinous hooks were usually no bigger than , though a belemnite could have had between 100 and 800 hooks in total, using them to stab and hold onto prey. Belemnites were an important food source for many Mesozoic marine creatures, both the adults and the planktonic juveniles and they likely played an important role in restructuring marine ecosystems after the Triassic–Jurassic extinction event. They may have laid between 100 and 1,000 eggs. Some species ...
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Per Mille
The phrase per mille () indicates parts per thousand. The associated symbol is , similar to a per cent sign but with an extra zero in the division (mathematics), divisor. Major dictionaries do not agree on the spelling, giving other options of per mil, per mill, permil, permill, permille. The word is the cognate in Dutch, German, Finnish and Swedish, and is sometimes seen as a loanword in English with the same meaning as ''per mille''. (Page 258) The symbol is included in the General Punctuation (Unicode block), General Punctuation block of Unicode at .Unicode.General Punctuation. 2014. Accessed 5 Aug 2014. There is also an Arabic-Indic per mille sign at . Examples Blood alcohol concentration Promille and permille are likely best known as shorthand terms for Blood Alcohol Concentration, where 1 promille corresponds to 1 gram of alcohol per liter of blood. Promille is used in the legal limits of BAC by volume, blood-alcohol content for driving a road vehicle in some count ...
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Δ13C
In geochemistry, paleoclimatology, and paleoceanography ''δ''13C (pronounced "delta thirteen c") is an isotopic signature, a measure of the ratio of the two stable isotopes of carbon—Carbon-13, 13C and Carbon-12, 12C—reported in parts per thousand (per mil, ‰). The measure is also widely used in archaeology for the reconstruction of past diets, particularly to see if marine foods or certain types of plants were consumed. The definition is, in per mille: :\delta \ce = \left( \frac - 1 \right) \times 1000 where the standard is an established Reference materials for stable isotope analysis, reference material. ''δ''13C varies in time as a function of productivity, the signature of the inorganic source, Oceanic carbon cycle#Burial, organic carbon burial, and vegetation type. Biological processes preferentially take up the lower mass isotope through kinetic fractionation. However some abiotic processes do the same. For example, methane from hydrothermal vents can be depleted ...
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Isotope Analysis
Isotope analysis is the identification of isotopic signature, abundance of certain stable isotopes of chemical elements within organic and inorganic compounds. Isotopic analysis can be used to understand the flow of energy through a food web, to reconstruct past environmental and climatic conditions, to investigate human and animal diets, for food authentification, and a variety of other physical, geological, palaeontological and chemical processes. Stable isotope ratios are measured using mass spectrometry, which separates the different isotopes of an element on the basis of their mass-to-charge ratio. Tissues affected Isotopic oxygen is incorporated into the body primarily through ingestion at which point it is used in the formation of, for archaeological purposes, bones and teeth. The oxygen is incorporated into the hydroxylcarbonic apatite of bone and tooth enamel. Bone is continually remodelled throughout the lifetime of an individual. Although the rate of tur ...
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Isotope Geochemistry
Isotope geochemistry is an aspect of geology based upon the study of natural variations in the relative abundances of isotopes of various Chemical element, elements. Variations in isotopic abundance are measured by isotope-ratio mass spectrometry, and can reveal information about the ages and origins of rock, air or water bodies, or processes of mixing between them. Stable isotope geochemistry is largely concerned with isotopic variations arising from mass-dependent isotope fractionation, whereas radiogenic isotope geochemistry is concerned with the products of natural radioactivity. Stable isotope geochemistry For most stable isotopes, the magnitude of fractionation from kinetic fractionation, kinetic and equilibrium fractionation is very small; for this reason, enrichments are typically reported in "per mil" (Per mille, ‰, parts per thousand). These enrichments (δ) represent the ratio of heavy isotope to light isotope in the sample over the ratio of a Reference Materials fo ...
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Carbon-12
Carbon-12 (12C) is the most abundant of the two stable isotopes of carbon ( carbon-13 being the other), amounting to 98.93% of element carbon on Earth; its abundance is due to the triple-alpha process by which it is created in stars. Carbon-12 is of particular importance in its use as the standard from which atomic masses of all nuclides are measured, thus, its atomic mass is exactly 12 daltons by definition. Carbon-12 is composed of 6 protons, 6 neutrons, and 6 electrons. History Before 1959, both the IUPAP and IUPAC used oxygen to define the mole; the chemists defining the mole as the number of atoms of oxygen which had mass 16 g, the physicists using a similar definition but with the oxygen-16 isotope only. The two organizations agreed in 1959–60 to define the mole as follows. ''Mole is the amount of substance of a system which contains as many elementary entities as there are atoms in 12 gram of carbon 12; its symbol is "mol".'' This was adopted by the ...
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Figure 14 - With This Method C3, C4 And CAM Plant Metabolism Are Well Separated
Figure may refer to: General *A shape, drawing, depiction, or geometric configuration *Figure (wood), wood appearance *Figure (music), distinguished from musical motif *Noise figure, in telecommunication *Dance figure, an elementary dance pattern *A person's figure, human physical appearance *Figure–ground (perception), the distinction between a visually perceived object and its surroundings Arts *Figurine, a miniature statuette representation of a creature *Action figure, a posable jointed solid plastic character figurine *Figure painting, realistic representation, especially of the human form *Figure drawing *Model figure, a scale model of a creature Writing *figure, in writing, a type of floating block (text, table, or graphic separate from the main text) *Figure of speech, also called a rhetorical figure *Christ figure, a type of character * in typesetting, text figures and lining figures Accounting *Figure, a synonym for number *Significant figures in a decimal number ...
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