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Anomalocarids
Radiodonta is an extinct Order (biology), order of stem-group arthropods that was successful worldwide during the Cambrian period. Radiodonts are distinguished by their distinctive frontal appendages, which are morphologically diverse and were used for a variety of functions. Radiodonts were among the earliest large predators, but they also included sediment sifters and filter feeders. Some of the most famous species of radiodonts are the Cambrian taxa ''Anomalocaris, Anomalocaris canadensis'', ''Hurdia, Hurdia victoria'', ''Peytoia nathorsti'', ''Titanokorys gainesii, Titanokorys gainesi, Cambroraster, Cambroraster falcatus'' and ''Amplectobelua, Amplectobelua symbrachiata''. The later surviving members include the subfamily Aegirocassisinae from the Early Ordovician of Morocco and the Early Devonian member ''Schinderhannes bartelsi'' from Germany. Etymology The name Radiodonta (Latin for ''radius'' "spoke of a wheel" and Greek for ''odoús'' "tooth") refers to the radial arrang ...
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Arthropods
Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metamerism (biology), metameric) Segmentation (biology), segments, and paired jointed appendages. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. They form an extremely diverse group of up to ten million species. Haemolymph is the analogue of blood for most arthropods. An arthropod has an open circulatory system, with a body cavity called a haemocoel through which haemolymph circulates to the interior Organ (anatomy), organs. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. They have ladder-like nervous systems, with paired Anatomical terms of location#Dorsal and ventral, ventral Ventral nerve cord, nerve cord ...
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Laminacaris
''Laminacaris'' is a genus of extinct stem-group arthropods (Radiodonta) that lived during the Cambrian period. It is monotypic with a single species ''Laminacaris chimera'', the fossil of which was described from the Chengjiang biota of China in 2018. Around the same time, two specimens that were similar or of the same species were discovered at the Kinzers Formation in Pennsylvania, USA. The first specimens from China were three frontal appendages, without the other body parts. Discovery and naming ''Laminacaris'' specimens were discovered from the mudstones of the Yu'anshan Member of the Chiungchussu Formation in eastern Yunnan Province, the Mafang section in the Haikou area at Kunming, and the Fengkoushao section , Chengjiang. The name is derived from Latin words, ''lamina'' meaning thin blade, and ''caris'' for crab; the species name refers to a Greek mythological creature, ''Chimera'', that has a body composed of parts of more than one animal. Description The large fro ...
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Paranomalocaris
''Paranomalocaris'' is a genus of radiodont recovered from the Cambrian Stage 4 aged Wulongqing Formation, eastern Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces .... It contains two species, ''Paranomalocaris multisegmentalis'' and ''P. simplex''. It is only known from its frontal appendage. The frontal appendage features endites with numerous auxiliary spines, and at least 22 segments (podomeres), hence the specific name ''multisegmentalis''. References {{Taxonbar, from=Q65042329 Radiodonta Fossil taxa described in 2013 Cambrian arthropods ...
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Tagma (biology)
In biology, a tagma (Greek: τάγμα, : tagmata – τάγματα - body of soldiers; battalion) is a specialized grouping of multiple segmentation (biology), segments or Metamerism (biology), metameres into a coherently functional Morphology (biology), morphological unit. Familiar examples are the head, the Thorax (arthropod anatomy), thorax, and the Abdomen#Arthropoda, abdomen of insects. The segments within a tagma may be either fused (such as in the head of an insect) or so jointed as to be independently moveable (such as in the abdomen of most insects). Usually the term is taken to refer to tagmata in the morphology of members of the phylum Arthropoda, but it applies equally validly in other phyla, such as the Chordata. In a given taxon the names assigned to particular tagmata are in some sense informal and arbitrary; for example, not all the tagmata of species within a given subphylum of the Arthropoda are Homology (biology), homologous to those of species in other subph ...
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Schinderhannes Bartelsi
''Schinderhannes bartelsi'' is a fossil species of hurdiid radiodont (anomalocaridid), known from a single specimen from the Lower Devonian Hunsrück Slate in Germany. Its discovery expanded the known range of radiodonts, the latest members of which were previously known only from the Early Ordovician, at least 66 million years earlier than ''Schinderhannes bartelsi''. Discovery The single specimen was discovered in the Eschenbach-Bocksberg Quarry in Bundenbach, Rhineland-Palatinate, Germany, and is named after the outlaw Schinderhannes who frequented the area. Its specific epithet ''bartelsi'' honours Christoph Bartels, a Hunsrück Slate expert. The specimen is now housed in the Naturhistorisches Museum in Mainz. Morphology ''Schinderhannes'' is about long in full body length ( long excluding telsonDryad Data
). Like ...
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Early Devonian
The Early Devonian is the first of three Epoch (geology), epochs comprising the Devonian period, corresponding to the Lower Devonian Series (stratigraphy), series. It lasted from and began with the Lochkovian Stage , which was followed by the Pragian from and then by the Emsian, which lasted until the Middle Devonian began, . During this time, the first Ammonoidea, ammonoids appeared, descending from Bactritida, bactritoid Nautiloidea, nautiloids. Ammonoids during this time period were simple and differed little from their nautiloid counterparts. These ammonoids belong to the order Agoniatitida, which in later epochs evolved to new ammonoid orders, for example Goniatite, Goniatitida and Clymeniida. This class of cephalopod molluscs would dominate the marine fauna until the beginning of the Mesozoic Era. References

Early Devonian, Geological epochs Devonian geochronology, *01 {{Geochronology-stub ...
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Morocco
Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocco border, the east, and the disputed territory of Western Sahara to Morocco–Western Sahara border, the south. Morocco also claims the Spain, Spanish Enclave and exclave, exclaves of Ceuta, Melilla and Peñón de Vélez de la Gomera, and several small Plazas de soberanía, Spanish-controlled islands off its coast. It has a population of approximately 37 million. Islam is both the official and predominant religion, while Arabic and Berber are the official languages. Additionally, French and the Moroccan dialect of Arabic are widely spoken. The culture of Morocco is a mix of Arab culture, Arab, Berbers, Berber, Culture of Africa, African and Culture of Europe, European cultures. Its capital is Rabat, while its largest city is Casablanca. Th ...
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Early Ordovician
The Early Ordovician is the first epoch (geology), epoch of the Ordovician period, corresponding to the Lower Ordovician series (stratigraphy), series of the Ordovician system. It began after the Cambrian Stage 10, Age 10 of the Furongian epoch of the Cambrian and lasted from to million years ago, until the Dapingian age of the Middle Ordovician. It includes Tremadocian and Floian ages. History International Commission on Stratigraphy (ICS) appointed working groups on the boundaries of the Ordovician subdivisions in 1974. The boundaries were established by the fauna of conodonts and/or graptolites. In 1995, the Subcommission on Ordovician Stratigraphy, with the support of 90% majority of voting members, adopted the division of the Ordovician system into three series: Lower, Middle Ordovician, Middle and Upper Ordovician, Upper. In the same year, it was decided to divide each of the three series into two global stages. ''Tetragraptus approximatus'' zone was chosen as the base ...
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Aegirocassisinae
Aegirocassisinae is a subfamily of radiodonts (marine stem-arthropods) from the lower Paleozoic era. It belongs to the larger hurdiidae (peytoiid) family, which were the most diverse and long lasting radiodonts. The members of this subfamily are restricted to the Lower Ordovician-aged Fezouata Formation of Morocco. Currently only two genera are included: ''Aegirocassis'' and '' Pseudoangustidontus''. These two genera possess large Baleen-like auxiliary spines on their frontal appendages, which suggests a suspension feeding lifestyle for the group. These radiodonts are some of the few known from sediments beyond the Cambrian period. This subfamily shows that following the Great Ordovician Biodiversification Event, which saw a rise in the plankton population in the worlds oceans, suspension feeding became more common in radiodonts then other feeding styles. It also seems that due to the evolution of new predators, like large nautiloid cephalopods, and other arthropod groups like the ...
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Titanokorys Gainesii
''Titanokorys'' is a genus of extinct hurdiid (peytoiid) radiodont (a grouping of primitive stem arthropods which lived during the early Paleozoic) that lived during the Miaolingian epoch of the middle Cambrian. It is the largest member of its family from the Cambrian, with an estimated body length of around long, making it one of the largest animals of the time. It bears a resemblance to the related, and contemporary, genus '' Cambroraster''. Fossils of ''T. gainesi'' were first found within the Marble Canyon locality of the Burgess Shale in 2018, however the fossils were not named until 2021 because they were assumed to be giant specimens of ''Cambroraster''. The taxon is one of several genera of radiodonts known from the Burgess Shale, with some of the others being ''Cambroraster'', ''Anomalocaris'', ''Peytoia'', and '' Hurdia''. ''Titanokorys'' is distinguished from other Burgess Shale radiodonts because of its large anterior sclerite (head covering carapace, or H-eleme ...
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Cambrian
The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordovician Period 486.85 Ma. Most of the continents lay in the southern hemisphere surrounded by the vast Panthalassa Ocean. The assembly of Gondwana during the Ediacaran and early Cambrian led to the development of new convergent plate boundaries and continental-margin arc magmatism along its margins that helped drive up global temperatures. Laurentia lay across the equator, separated from Gondwana by the opening Iapetus Ocean. The Cambrian marked a profound change in life on Earth; prior to the Period, the majority of living organisms were small, unicellular and poorly preserved. Complex, multicellular organisms gradually became more common during the Ediacaran, but it was not until the Cambrian that fossil diversity seems to rapidly ...
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Stem-group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown ...
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