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Amoeboflagellates
An amoeboflagellate ( amoeboflagellates) is any eukaryotic organism capable of behaving as an amoeba and as a flagellate at some point during their life cycle. Amoeboflagellates present both pseudopodia and at least one flagellum, often simultaneously. Occurrence The amoeboflagellate cell type has been acquired numerous independent times across the evolution of protists (i.e. primarily unicellular eukaryotes that are not plants, fungi or animals). Some examples of protist phyla with amoeboflagellate body types are: * Cercozoa contains various examples of amoeboflagellates with filose pseudopods, thread-like cell projections also known as filopodia. The cercomonads, glissomonads and paracercomonads behave as amoeboflagellates with two flagella throughout the majority of their life cycle, and are essential predators of the soil microbiome. Among the more basal Cercozoa is '' Reticulamoeba'', an amoeboflagellate with reticulose pseudopods (reticulopodia), filopodia that join ...
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Filosa
Cercozoa (now synonymised with Filosa) is a phylum of diverse single-celled eukaryotes. They lack shared morphological characteristics at the microscopic level, and are instead united by phylogeny, molecular phylogenies of rRNA and actin or Ubiquitin#Polyubiquitin chains, polyubiquitin. They were the first major eukaryotic group to be recognized mainly through phylogeny, molecular phylogenies. They are the natural predators of many species of bacteria. They are closely related to the phylum Retaria, comprising amoeboids that usually have complex shells, and together form a supergroup called Rhizaria. Characteristics The group includes most amoeboids and flagellates that feed by means of filose pseudopods. These may be restricted to part of the cell surface, but there is never a true cytostome or mouth as found in many other protozoa. They show a variety of forms and have proven difficult to define in terms of structural characteristics, although their unity is strongly supported b ...
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Protist
A protist ( ) or protoctist is any eukaryotic organism that is not an animal, land plant, or fungus. Protists do not form a natural group, or clade, but are a paraphyletic grouping of all descendants of the last eukaryotic common ancestor excluding land plants, animals, and fungi. Protists were historically regarded as a separate taxonomic kingdom known as Protista or Protoctista. With the advent of phylogenetic analysis and electron microscopy studies, the use of Protista as a formal taxon was gradually abandoned. In modern classifications, protists are spread across several eukaryotic clades called supergroups, such as Archaeplastida ( photoautotrophs that includes land plants), SAR, Obazoa (which includes fungi and animals), Amoebozoa and " Excavata". Protists represent an extremely large genetic and ecological diversity in all environments, including extreme habitats. Their diversity, larger than for all other eukaryotes, has only been discovered in rece ...
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Cercozoa
Cercozoa (now synonymised with Filosa) is a phylum of diverse single-celled eukaryotes. They lack shared morphological characteristics at the microscopic level, and are instead united by phylogeny, molecular phylogenies of rRNA and actin or Ubiquitin#Polyubiquitin chains, polyubiquitin. They were the first major eukaryotic group to be recognized mainly through phylogeny, molecular phylogenies. They are the natural predators of many species of bacteria. They are closely related to the phylum Retaria, comprising amoeboids that usually have complex shells, and together form a supergroup called Rhizaria. Characteristics The group includes most amoeboids and flagellates that feed by means of filose pseudopods. These may be restricted to part of the cell surface, but there is never a true cytostome or mouth as found in many other protozoa. They show a variety of forms and have proven difficult to define in terms of structural characteristics, although their unity is strongly supported b ...
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Amoeba
An amoeba (; less commonly spelled ameba or amœba; : amoebas (less commonly, amebas) or amoebae (amebae) ), often called an amoeboid, is a type of Cell (biology), cell or unicellular organism with the ability to alter its shape, primarily by extending and retracting pseudopodia, pseudopods. Amoebae do not form a single Taxonomy (biology), taxonomic group; instead, they are found in every major Lineage (evolution), lineage of eukaryote, eukaryotic organisms. Amoeboid cells occur not only among the protozoa, but also in fungi, algae, and animals. Microbiologists often use the terms "amoeboid" and "amoeba" interchangeably for any organism that exhibits amoeboid movement. In older classification systems, most amoebae were placed in the Class (biology), class or subphylum Sarcodina, a grouping of Unicellular organism, single-celled organisms that possess pseudopods or move by protoplasmic flow. However, molecular phylogenetic studies have shown that Sarcodina is not a monophyletic ...
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Flagellate
A flagellate is a cell or organism with one or more whip-like appendages called flagella. The word ''flagellate'' also describes a particular construction (or level of organization) characteristic of many prokaryotes and eukaryotes and their means of motion. The term presently does not imply any specific relationship or classification of the organisms that possess flagella. However, several derivations of the term "flagellate" (such as " dinoflagellate" and " choanoflagellate") are more formally characterized. Form and behavior Flagella in eukaryotes are supported by microtubules in a characteristic arrangement, with nine fused pairs surrounding two central singlets. These arise from a basal body. In some flagellates, flagella direct food into a cytostome or mouth, where food is ingested. Flagella role in classifying eukaryotes. Among protoctists and microscopic animals, a flagellate is an organism with one or more flagella. Some cells in other animals may be flage ...
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Naegleria Fowleri Lifecycle Stages
''Naegleria'' is a genus consisting of 47 described species of protozoa often found in warm aquatic environments as well as soil habitats worldwide. It has three life cycle forms: the amoeboid stage, the cyst stage, and the flagellated stage, and has been routinely studied for its ease in change from amoeboid to flagellated stages. The ''Naegleria'' genera became famous when ''Naegleria fowleri'', the causative agent of the usually fatal human and animal disease primary amoebic meningoencephalitis (PAM), was discovered in 1965. Most species in the genus, however, are incapable of causing disease. Etymology The genus ''Naegleria'' is named after the German protozoologist, Kurt Nägler. History In 1899, Franz Schardinger discovered an amoeba that had the ability to transform into a flagellated stage. He named the organism ''Amoeba gruberi'', which was later changed to the genus ''Naegleria'' in 1912 by Alexeieff. Before 1970, the genus was generally used as a model organism ...
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Amoebozoa
Amoebozoa is a major Taxonomy (biology), taxonomic group containing about 2,400 described species of Amoeba, amoeboid protists, often possessing blunt, fingerlike, Pseudopod#Morphology, lobose pseudopods and tubular mitochondrial cristae. In traditional classification schemes, Amoebozoa is usually ranked as a phylum within either the kingdom (biology), kingdom Protista or the kingdom Protozoa. In the classification favored by the International Society of Protistologists, it is retained as an unranked "supergroup (biology), supergroup" within Eukaryota. Molecular genetics, Molecular genetic analysis supports Amoebozoa as a monophyletic clade. Modern studies of eukaryotic phylogenetic trees identify it as the sister group to Opisthokonta, another major clade which contains both fungi and animals as well as several other clades comprising some 300 species of unicellular eukaryotes. Amoebozoa and Opisthokonta are sometimes grouped together in a high-level taxon, named Amorphea. Amoeboz ...
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Soil Microbiome
Soil microbiology is the study of microorganisms in soil, their functions, and how they affect soil properties. It is believed that between two and four billion years ago, the first ancient bacteria and microorganisms came about on Earth's oceans. These bacteria could fix nitrogen, in time multiplied, and as a result released oxygen into the atmosphere. This led to more advanced microorganisms, which are important because they affect soil structure and fertility. Soil microorganisms can be classified as bacteria, actinomycetes, fungi, algae and protozoa. Each of these groups has characteristics that define them and their functions in soil.Rao, Subba. Soil Microbiology. Fourth ed. Enfield: Science Publishers, 1999. Print. Up to 10 billion bacterial cells inhabit each gram of soil in and around plant roots, a region known as the rhizosphere. In 2011, a team detected more than 33,000 bacterial and archaeal species on sugar beet roots. The composition of the rhizobiome can change rap ...
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Reticulamoeba
''Reticulamoeba'' is a genus of Rhizaria. It is a relatively large unicellular protist and is commonly known for its thin reticulopodia radiating outwards from the cell (Bass et al., 2012). ''Reticulamoeba'' are strictly benthic protists that use their reticulopodia for feeding. They have a close association with benthic diatoms as their main source of food (Bass et al., 2012). There are only two ''Reticulamoeba'' species described to date and the protozoologist Karl Grell discovered both of them (Bass et al., 2012). History The genus ''Reticulamoeba'' was described in 1994 by the protozoologist Karl Grell based on the species R. gemmipara and not until a year later a second species R. minor was discovered (Bass et al., 2012). The two species form a clade within the cercozoan class Granofilosea (Bass et al., 2012). Within this clade, ''Reticulamoeba'' present phylogenetic similarities with the genus Mesofila (Bass et al., 2012). Habitat and ecology ''Reticulamoeba'' is common ...
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Myxogastrid
Myxogastria/Myxogastrea (myxogastrids, ICZN) or Myxomycetes ( ICN) is a class of slime molds that contains 5 orders, 14 families, 62 genera, and 888 species. They are colloquially known as the ''plasmodial'' or ''acellular'' slime moulds. All species pass through several very different morphologic phases, such as microscopic individual cells, slimy amorphous organisms visible with the naked eye, and conspicuously shaped fruit bodies. Although they are monocellular, they can reach immense widths and weights: in extreme cases they can be up to across and weigh up to . The class Myxogastria is distributed worldwide, but it is more common in temperate regions where it has a higher biodiversity than in polar regions, the subtropics, or the tropics. They are mainly found in open forests, but also in extreme regions such as deserts, under snow blankets, or underwater. They also occur on the bark of trees, sometimes high in the canopy. These are known as corticol ...
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Lobosa
Lobosa is a taxonomic group of amoebae in the phylum Amoebozoa. Most lobosans possess broad, bluntly rounded pseudopods, although one genus in the group, the recently discovered ''Sapocribrum,'' has slender and threadlike (filose) pseudopodia. In current classification schemes, Lobosa is a subphylum, composed mainly of amoebae that have lobose pseudopods but lack cilia or flagella. Characteristics The group was originally proposed in 1861 by William B. Carpenter, who created it as a taxonomic order containing the single family Amoebina. Carpenter's Lobosa consisted of amoeboid organisms whose endoplasm (endosarc) flows into lobe-like "pseudopodian prolongations." This type of pseudopod, which was understood to be typical of the genus Amoeba "and its allies," differed from the filose (thread-like) or reticulose (netlike) pseudopods of the Foraminifera. The name Lobosa was chosen for these amoebae "as expressing the lobe-like character of their pseudopodial extensions". As cur ...
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