Zapfella
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Zapfella
''Rogerella'' is a small pouch-shaped boring (a type of trace fossil) with a slit-like aperture currently produced by acrothoracican barnacles. These crustaceans extrude their legs upwards through the opening for filter-feeding. They are known in the fossil record as borings in carbonate substrates (shells and hardgrounds) from the Devonian The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian per ... to the Recent. References {{Taxonbar, from=Q7359233 Boring fossils Ichnotaxa ...
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Bioerosion
Bioerosion describes the breakdown of hard ocean substrates – and less often terrestrial substrates – by living organisms. Marine bioerosion can be caused by mollusks, polychaete worms, phoronids, sponges, crustaceans, echinoids, and fish; it can occur on coastlines, on coral reefs, and on ships; its mechanisms include biotic boring, drilling, rasping, and scraping. On dry land, bioerosion is typically performed by pioneer plants or plant-like organisms such as lichen, and mostly chemical (e.g. by acidic secretions on limestone) or mechanical (e.g. by roots growing into cracks) in nature. Bioerosion of coral reefs generates the fine and white coral sand characteristic of tropical islands. The coral is converted to sand by internal bioeroders such as algae, fungi, bacteria (microborers) and sponges (Clionaidae), bivalves (including '' Lithophaga''), sipunculans, polychaetes, acrothoracican barnacles and phoronids, generating extremely fine sediment with diame ...
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Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. The Jurassic constitutes the second and middle period of the Mesozoic, Mesozoic Era as well as the eighth period of the Phanerozoic, Phanerozoic Eon and is named after the Jura Mountains, where limestone strata from the period were first identified. The start of the Jurassic was marked by the major Triassic–Jurassic extinction event, associated with the eruption of the Central Atlantic magmatic province, Central Atlantic Magmatic Province (CAMP). The beginning of the Toarcian Age started around 183 million years ago and is marked by the Toarcian Oceanic Anoxic Event, a global episode of Anoxic event, oceanic anoxia, ocean acidification, and elevated global temperatures associated with extinctions, likely caused by the eruption of the Kar ...
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Callovian
In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma (million years ago) and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the Oxfordian. Stratigraphic definitions The Callovian Stage was first described by French palaeontologist Alcide d'Orbigny in 1852. Its name derives from the latinized name for Kellaways Bridge, a small hamlet 3 km north-east of Chippenham, Wiltshire, England. The base of the Callovian is defined as the place in the stratigraphic column where the ammonite genus '' Kepplerites'' first appears, which is the base of the biozone of '' Macrocephalites herveyi''. A global reference profile (a GSSP) for the base had in 2009 not yet been assigned. The top of the Callovian (the base of the Oxfordian) is at the first appearance of ammonite species '' Brightia thuouxensis''. Subdivision The Callovian is often subdivided into three substages ...
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Crinoid
Crinoids are marine invertebrates that make up the class Crinoidea. Crinoids that remain attached to the sea floor by a stalk in their adult form are commonly called sea lilies, while the unstalked forms, called feather stars or comatulids, are members of the largest crinoid order, Comatulida. Crinoids are echinoderms in the phylum Echinodermata, which also includes the starfish, brittle stars, sea urchins and sea cucumbers. They live in both shallow water and in depths of over . Adult crinoids are characterised by having the mouth located on the upper surface. This is surrounded by feeding arms, and is linked to a U-shaped gut, with the anus being located on the oral disc near the mouth. Although the basic echinoderm pattern of fivefold symmetry can be recognised, in most crinoids the five arms are subdivided into ten or more. These have feathery pinnules and are spread wide to gather planktonic particles from the water. At some stage in their lives, most crinoids have a sho ...
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Matmor Formation
The Matmor Formation is a geologic formation of up to thick, that is exposed in Hamakhtesh Hagadol in southern Israel. The Matmor Formation contains fossils from a Jurassic equatorial shallow marine environment. Bivalves, gastropods, sponges, corals, echinoderms, and sclerobionts are present in the Matmor Formation to various degrees (Wilson et al., 2010). The stratigraphy of the Matmor Formation consists of alternating layers of limestone and marl (Hirsch and Roded, 1996). Stratigraphy The Iraq Petroleum Company originally described the Jurassic sequence in Hamakhtesh Hagadol of Israel in the 1930s (Hudson, 1958). These findings were later described and published by Blake (1935). Shaw (1947) published a limited summary of the stratigraphy. Hudson (1958) later subdivided the rocks into the Callovian, Divesian, Argovian, Sequanim biostratigraphic stages. In 1963, Goldberg subdivided the section into the Zohar, Kidod and Be'er Sheva Formations. Goldberg (1963) further divide ...
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Israel
Israel, officially the State of Israel, is a country in West Asia. It Borders of Israel, shares borders with Lebanon to the north, Syria to the north-east, Jordan to the east, Egypt to the south-west, and the Mediterranean Sea to the west. Israeli-occupied territories, It occupies the Occupied Palestinian territories, Palestinian territories of the West Bank in the east and the Gaza Strip in the south-west. Israel also has a small coastline on the Red Sea at its southernmost point, and part of the Dead Sea lies along its eastern border. Status of Jerusalem, Its proclaimed capital is Jerusalem, while Tel Aviv is the country's Gush Dan, largest urban area and Economy of Israel, economic center. Israel is located in a region known as the Land of Israel, synonymous with the Palestine (region), Palestine region, the Holy Land, and Canaan. In antiquity, it was home to the Canaanite civilisation followed by the History of ancient Israel and Judah, kingdoms of Israel and Judah. Situate ...
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Trace Fossil
A trace fossil, also called an ichnofossil (; ), is a fossil record of biological activity by lifeforms, but not the preserved remains of the organism itself. Trace fossils contrast with body fossils, which are the fossilized remains of parts of organisms' bodies, usually altered by later chemical activity or by mineralization. The study of such trace fossils is ichnology - the work of ichnologists. Trace fossils may consist of physical impressions made on or in the substrate by an organism. For example, burrows, borings ( bioerosion), urolites (erosion caused by evacuation of liquid wastes), footprints, feeding marks, and root cavities may all be trace fossils. The term in its broadest sense also includes the remains of other organic material produced by an organism; for example coprolites (fossilized droppings) or chemical markers (sedimentological structures produced by biological means; for example, the formation of stromatolites). However, most sedimentary struct ...
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Acrothoracica
The Acrothoracica are an infraclass of barnacles. Acrothoracicans bore into calcareous material such as mollusc shells, coral, crinoids or hardgrounds, producing a slit-like hole in the surface known by the trace fossil name '' Rogerella''. Acrothoracicans are typically smaller than other types of barnacle, being only a few millimetres in length. Being protected by the hard surfaces into which they have bored, they have no solid carapace of plates like other barnacles but have a soft, sac-like body fixed to the surface by a chitinous disc at the front of the head. They have from four to six pairs of feathery limbs, or "cirri", which they project out of their borings to catch drifting detritus for food. The mouthparts consist of mandibles, maxillules and maxillae. One pair of cirri is close to these while the others are at the other end of the body. The cyprid larva in Lithoglyptidae and Trypetesidae have a well-developed carapace which can completely enclose the body, a dev ...
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Barnacles
Barnacles are arthropods of the subclass Cirripedia in the subphylum Crustacea. They are related to crabs and lobsters, with similar nauplius larvae. Barnacles are exclusively marine invertebrates; many species live in shallow and tidal waters. Some 2,100 species have been described. Barnacle adults are sessile; most are suspension feeders with hard calcareous shells, but the Rhizocephala are specialized parasites of other crustaceans, with reduced bodies. Barnacles have existed since at least the mid-Carboniferous, some 325 million years ago. In folklore, barnacle geese were once held to emerge fully formed from goose barnacles. Both goose barnacles and the Chilean giant barnacle are fished and eaten. Barnacles are economically significant as biofouling on ships, where they cause hydrodynamic drag, reducing efficiency. Etymology The word "barnacle" is attested in the early 13th century as Middle English "bernekke" or "bernake", close to Old French "bernaque" and med ...
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Filter Feeder
Filter feeders are aquatic animals that acquire nutrients by feeding on organic matters, food particles or smaller organisms (bacteria, microalgae and zooplanktons) suspended in water, typically by having the water pass over or through a specialized filtering organ that sieves out and/or traps solids. Filter feeders can play an important role in condensing biomass and removing excess nutrients (such as nitrogen and phosphate) from the local waterbody, and are therefore considered water-cleaning ecosystem engineers. They are also important in bioaccumulation and, as a result, as indicator organisms. Filter feeders can be sessile, planktonic, nektonic or even neustonic (in the case of the buoy barnacle) depending on the species and the niches they have evolved to occupy. Extant species that rely on such method of feeding encompass numerous phyla, including poriferans ( sponges), cnidarians (jellyfish, sea pens and corals), arthropods ( krill, mysids and barna ...
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Hardgrounds
Carbonate hardgrounds are surfaces of synsedimentarily cemented carbonate layers that have been exposed on the seafloor (Wilson and Palmer, 1992). A hardground is essentially, then, a lithified seafloor. Ancient hardgrounds are found in limestone sequences and distinguished from later-lithified sediments by evidence of exposure to normal marine waters. This evidence can consist of encrusting marine organisms (especially bryozoans, oysters, barnacles, cornulitids, hederelloids, microconchids and crinoids), borings of organisms produced through bioerosion, early marine calcite cements, or extensive surfaces mineralized by iron oxides or calcium phosphates (Palmer, 1982; Bodenbender et al., 1989; Vinn and Wilson, 2010; Vinn and Toom, 2015). Modern hardgrounds are usually detected by sounding in shallow water or through remote sensing techniques like side-scan sonar. Carbonate hardgrounds often host a unique fauna and flora adapted to the hard surface. Organisms usually cemen ...
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Devonian
The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago (Megaannum, Ma), to the beginning of the succeeding Carboniferous period at Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied. The first significant evolutionary radiation of history of life#Colonization of land, life on land occurred during the Devonian, as free-spore, sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaf, leaves and true roots, and by the end of the period the first seed-bearing plants (Pteridospermatophyta, pteridospermatophyt ...
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