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Variable Checkerspot
The variable checkerspot or Chalcedon checkerspot (''Euphydryas chalcedona'') is a butterfly in the family Nymphalidae. It is found in western North America, where its range stretches from Alaska in the north to Baja California in the south and extends east through the Rocky Mountains into Colorado, Montana, New Mexico and Wyoming. The butterfly is usually brown or black with extensive white and yellow checkering and some red coloration on the dorsal wing. Adult wingspan is . Adult butterflies feed on nectar from flowers while larvae feed on a variety of plants including snowberry (''Symphoricarpos''), paintbrush (''Castilleja''), ''Buddleja'', ''Diplacus aurantiacus'' and ''Scrophularia californica''. During the breeding period, males congregate around larval host plants to encounter females. Males court female butterflies via physical displays.Rutowski, Ronald L., and George W. Gilchrist. "Courtship, Copulation and Oviposition in the Chalcedon Checkerspot,(Lepidoptera: Nymphalid ...
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William O
William is a masculine given name of Germanic origin. It became popular in England after the Norman conquest in 1066,All Things William"Meaning & Origin of the Name"/ref> and remained so throughout the Middle Ages and into the modern era. It is sometimes abbreviated "Wm." Shortened familiar versions in English include Will or Wil, Wills, Willy, Willie, Bill, Billie, and Billy. A common Irish form is Liam. Scottish diminutives include Wull, Willie or Wullie (as in Oor Wullie). Female forms include Willa, Willemina, Wilma and Wilhelmina. Etymology William is related to the German given name ''Wilhelm''. Both ultimately descend from Proto-Germanic ''*Wiljahelmaz'', with a direct cognate also in the Old Norse name ''Vilhjalmr'' and a West Germanic borrowing into Medieval Latin ''Willelmus''. The Proto-Germanic name is a compound of *''wiljô'' "will, wish, desire" and *''helmaz'' "helm, helmet".Hanks, Hardcastle and Hodges, ''Oxford Dictionary of First Names'', Oxford ...
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Diplacus Aurantiacus
''Diplacus'' is a genus of flowering plants in the family Phrymaceae, which was traditionally placed in family Scrophulariaceae. It includes 49 species native to the western United States and northwestern Mexico. Most prefer dry and rocky areas. The genus ''Diplacus'' was first described by Thomas Nuttall in 1838. It was merged into ''Mimulus'' no later than 1905, until the 2012 restructuring. The 2012 restructuring of ''Mimulus'' by Barker, ''et al''., based largely upon DNA evidence, left seven species in ''Mimulus'', placed 111 into ''Erythranthe'' (species with axile placentation and long pedicels), placed 46 into ''Diplacus'' (species with parietal placentation and sessile flowers), placed two in '' Uvedalia'', and placed one each in '' Elacholoma'', '' Mimetanthe'', and '' Thyridia''. Species 49 species are accepted.
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Euphydryas
''Euphydryas'' is a genus of Nymphalidae butterflies. Species References Further reading * Glassberg, Jeffrey ''Butterflies through Binoculars: The West'' (2001) * Guppy, Crispin S. and Shepard, Jon H. ''Butterflies of British Columbia'' (2001) * James, David G. and Nunnallee, David ''Life Histories of Cascadia Butterflies'' (2011) * Pelham, Jonathan ''Catalogue of the Butterflies of the United States and Canada'' (2008) * Pyle, Robert Michael ''The Butterflies of Cascadia'' (2002) External links Butterflies and Moths of North AmericaButterflies of America
Euphydryas, Melitaeini Butterflies of North America Butterfly genera Taxa named by Samuel Hubbard Scudder {{Nymphalinae-stub ...
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Genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants of an ancestral taxon are grouped together (i.e. Phylogeneti ...
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Nymphalini
Nymphalini is a tribe (biology), tribe of nymphalid Nymphalidae, brush-footed butterflies. Common names include Admiral (other)#Other uses, admirals, anglewings, Polygonia, commas, and Aglais, tortoiseshells, but none of these is specific to one particular genus. The name anglewing butterflies is an English translation of a Latin term ''papiliones angulati'', [Michael Denis, Denis & Ignaz Schiffermüller, Schiffermüller], ([1775, 1776]). Based on an overall similarity in the angulate wing shape, a collective name: ''Papiliones angulati'' was employed for ''Papilio atalanta'', ''P. antiopa'', ''P. cardui'', ''P. c-album'', ''P. io'', ''P. polychloros'', ''P. urticae'', ''P. xanthomelas'', ''P. vaualbum'', ''P. levana'' and ''P. prorsa''. The term ''papiliones angulati'' is applied as a collective taxon name, which therefore needs no type species as specified in Article 42.3.1 of the International Code of Zoological Nomenclature. ''Papiliones angulati'' has thus been re-d ...
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Aglais
''Aglais'' is a Holarctic genus of Nymphalidae, brush-footed butterflies, containing the tortoiseshells. This genus is sometimes indicated as a subgenus of ''Nymphalis'' or simply being an unnecessary division from the genus ''Nymphalis'',Evans, Arthur V. (2008). ''National Wildlife Federation Field Guide to Insects and Spider of North America''. p. 293. Sterling Publishing, London. . which also includes tortoiseshells, but it is usually considered to be separate. This proposed separate genus is also considered "brushfooted butterflies" historically together with the other or separate ''Nymphalis'' species. Species References External links

* With images. {{Taxonbar, from=Q32469 Nymphalini Nymphalidae genera Taxa named by Johan Wilhelm Dalman ...
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Family (biology)
Family (, : ) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". The delineation of what constitutes a family—or whether a described family should be acknowledged—is established and decided upon by active taxonomists. There are not strict regulations for outlining or acknowledging a family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to a lack of widespread consensus within the scientific community ...
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Lepidoptera
Lepidoptera ( ) or lepidopterans is an order (biology), order of winged insects which includes butterflies and moths. About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organisms, making it the second largest insect order (behind Coleoptera) with 126 family (biology), families and 46 Taxonomic rank, superfamilies, and one of the most widespread and widely recognizable insect orders in the world. Lepidopteran species are characterized by more than three derived features. The most apparent is the presence of scale (anatomy), scales that cover the torso, bodies, large triangular Insect wing, wings, and a proboscis for siphoning nectars. The scales are modified, flattened "hairs", and give butterflies and moths their wide variety of colors and patterns. Almost all species have some form of membranous wings, except for a few that have reduced wings or are wingless. Mating and the laying of eggs is normally performe ...
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Order (biology)
Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. For some groups of organisms, their orders may follow consist ...
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Catalpol
Catalpol is an iridoid glucoside. This natural product falls in the class of iridoid glycosides, which are simply monoterpenes with a glucose molecule attached. Natural occurrence First isolated in 1962, catalpol was named for plants in the genus ''Catalpa'' in which it was discovered. Later in 1969, catalpol was found to be present in larger quantities in several plants in genus ''Rehmannia'' (Orobanchaceae). It is also found in plants belonging to several families, including, but not limited to, Scrophulariaceae, Lamiaceae (including scullcap), Plantaginaceae (''Plantago sp'') and Bignoniaceae, all of which being in the order Lamiales. Because they feed on these plants, variable checkerspot butterflies (''Euphydryas chalcedona'') contain high amounts of catalpol, which makes them unpalatable to predators and thus serves as a defense mechanism. Biosynthetic pathway Though first isolated in the 1960s, there has been very little investigation of the biosynthetic pathway o ...
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Pupation
A pupa (; : pupae) is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages thereof being egg, larva, pupa, and imago. The processes of entering and completing the pupal stage are controlled by the insect's hormones, especially juvenile hormone, prothoracicotropic hormone, and ecdysone. The act of becoming a pupa is called pupation, and the act of emerging from the pupal case is called eclosion or emergence. The pupae of different groups of insects have different names such as ''chrysalis'' for the pupae of butterflies and ''tumbler'' for those of the mosquito family. Pupae may further be enclosed in other structures such as cocoons, nests, or shells. Position in life cycle The pupal stage follows the larval stage, or in some cases a prepupal stage, and precedes adulthood ('' imago'') in insects with compl ...
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Diapause
In animal dormancy, diapause is the delay in development in response to regular and recurring periods of adverse environmental conditions.Tauber, M.J., Tauber, C.A., Masaki, S. (1986) ''Seasonal Adaptations of Insects''. Oxford University Press It is a physiological state with very specific initiating and inhibiting conditions. The mechanism is a means of surviving predictable, unfavorable environmental conditions, such as temperature extremes, drought, or reduced food availability. Diapause is observed in all the life stages of arthropods, especially insects. Activity levels of diapausing stages can vary considerably among species. Diapause may occur in a completely immobile stage, such as the pupae and eggs, or it may occur in very active stages that undergo extensive migrations, such as the adult monarch butterfly, ''Danaus plexippus''. In cases where the insect remains active, feeding is reduced and reproductive development is slowed or halted. Embryonic diapause, a somewhat ...
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