Tuckermannopsis Hengduanensis
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Tuckermannopsis Hengduanensis
''Tuckermannopsis'' is a genus of foliose lichens in the family Parmeliaceae. Taxonomy The genus was circumscribed in 1933 by Hungarian lichenologist Vilmos Kőfaragó-Gyelnik. The genus name ''Tuckermannopsis'' honours Edward Tuckerman (1817–1886), who was an American botanist and professor who made significant contributions to the study of lichens and other alpine plants. He was a founding member of the Natural History Society of Boston and most of his career was spent at Amherst College. '' Tuckermannopsis ciliaris'' was assigned as the type, and at that time, only species. The type species is a lichen, meaning it is erect, foliose, and with apothecia and pycnidia ( sexual and asexual reproductive structures, respectively) that are largely restricted to the margins of the lobes. Starting in the 1980s, the genus became a wastebasket taxon containing cetrarioid species of uncertain taxonomic affinities. In 2001, Ingvar Kärnefelt and Arne Thell attempted to delimit ''Tucker ...
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Wastebasket Taxon
Wastebasket taxon (also called a wastebin taxon, dustbin taxon or catch-all taxon) is a term used by some taxonomists to refer to a taxon that has the purpose of classifying organisms that do not fit anywhere else. They are typically defined by either their designated members' often superficial similarity to each other, or their ''lack'' of one or more distinct character states or by their ''not'' belonging to one or more other taxa. Wastebasket taxa are by definition either paraphyletic or polyphyletic, and are therefore not considered valid taxa under strict cladistic rules of taxonomy. The name of a wastebasket taxon may in some cases be retained as the designation of an evolutionary grade, however. Examples There are many examples of paraphyletic groups, but true "wastebasket" taxa are those that are known not to, and perhaps not intended to, represent natural groups, but are nevertheless used as convenient groups of organisms. The acritarchs are perhaps the most famous ex ...
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Ascospore
In fungi, an ascospore is the sexual spore formed inside an ascus—the sac-like cell that defines the division Ascomycota, the largest and most diverse Division (botany), division of fungi. After two parental cell nucleus, nuclei fuse, the ascus undergoes meiosis (halving of genetic material) followed by a mitosis (cell division), ordinarily producing eight genetically distinct haploid spores; most yeasts stop at four ascospores, whereas some moulds carry out extra post-meiotic divisions to yield dozens. Many asci build turgor, internal pressure and shoot their spores clear of the calm boundary layer, thin layer of still air enveloping the fruit body, whereas subterranean truffles depend on animals for biological dispersal, dispersal. Ontogeny, Development shapes both form and endurance of ascospores. A hook-shaped crozier aligns the paired nuclei; a double-biological membrane, membrane system then parcels each daughter nucleus, and successive wall layers of β-glucan, chitosan ...
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Ascus
An ascus (; : asci) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or species can occur in numbers of one (e.g. '' Monosporascus cannonballus''), two, four, or multiples of four. In a few cases, the ascospores can bud off conidia that may fill the asci (e.g. '' Tympanis'') with hundreds of conidia, or the ascospores may fragment, e.g. some '' Cordyceps'', also filling the asci with smaller cells. Ascospores are nonmotile, usually single celled, but not infrequently may be coenocytic (lacking a septum), and in some cases coenocytic in multiple planes. Mitotic divisions within the developing spores populate each resulting cell in septate ascospores with nuclei. The term ocular chamber, or oculus, refers to the epiplasm (the portion of cytoplasm not used in ascospore formation) that is surrounded by the ...
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Fruticose Lichen
A fruticose lichen is a form of lichen fungi that is characterized by a coral-like shrubby or bushy growth structure. It is formed from a symbiotic relationship of a photobiont such as green algae or less commonly cyanobacteria and one, two or more mycobionts. Fruticose lichens are not a monophyletic and holophyletic lineage, but are a form encountered in many classes. Fruticose lichens have a complex vegetation structure, and are characterized by an ascending, bushy or pendulous appearance. As with other lichens, many fruticose lichens can endure high degrees of desiccation. They grow slowly and often occur in habitats such as on tree barks, on rock surfaces and on soils in the Arctic and mountain regions. Characteristics Characteristic of fruticose lichen is the shape of the thallus. Like crustose lichen, fruticose lichen is composed of a holdfast which will act as an anchor for the lichen to grow in rock fissures, over loose sand or soil. Growth and structure Fruticose or ...
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Foliose Lichen
A foliose lichen is a lichen with flat, leaf-like , which are generally not firmly bonded to the substrate on which it grows. It is one of the three most common growth forms of lichens. It typically has distinct upper and lower surfaces, each of which is usually covered with a cortex; some, however, lack a lower cortex. The photobiont layer lies just below the upper cortex. Where present, the lower cortex is usually dark (sometimes even black), but occasionally white. Foliose lichens are attached to their substrate either by hyphae extending from the cortex or , or by root-like structures called . The latter, which are found only in foliose lichens, come in a variety of shapes, the specifics of which can aid in species identification. Some foliose lichens attach only at a single stout peg called a , typically located near the lichen's centre. Lichens with this structure are called "umbilicate". In general, medium to large epiphytic foliose lichens are moderately sensitive to air ...
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Thallus
Thallus (: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. A thallus usually names the entire body of a multicellular non-moving organism in which there is no organization of the tissues into organs. Many of these organisms were previously known as the thallophytes, a polyphyletic group of distantly related organisms. An organism or structure resembling a thallus is called thalloid, thalloidal, thalliform, thalline, or thallose. Even though thalli do not have organized and distinct parts ( leaves, roots, and stems) as do the vascular plants, they may have analogous structures that resemble their vascular "equivalents". The analogous structures have similar function or macroscopic structure, but different microscopic structure; for example, no thallus has vascular tissue. In exceptional cases such as the Lemnoideae, where th ...
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Systematics
Systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: phylogenetic trees, phylogenies). Phylogenies have two components: branching order (showing group relationships, graphically represented in cladograms) and branch length (showing amount of evolution). Phylogenetic trees of species and higher taxa are used to study the evolution of traits (e.g., anatomical or molecular characteristics) and the distribution of organisms ( biogeography). Systematics, in other words, is used to understand the evolutionary history of life on Earth. The word systematics is derived from the Latin word of Ancient Greek origin '' systema,'' which means systematic arrangement of organisms. Carl Linnaeus used 'Systema Naturae' as the title of his book. Branches and applications In the study of biological systematics, researchers use the different br ...
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Nephromopsis
''Nephromopsis'' is a genus of lichenized fungi within the Parmeliaceae The Parmeliaceae is a large and diverse family of Lecanoromycetes. With over 2700 species in 71 genera, it is the largest family of lichen-forming fungi. The most speciose genera in the family are the well-known groups: '' Xanthoparmelia'' ( 82 ... family. References External links Index Fungorum Parmeliaceae Lichen genera Lecanorales genera Taxa named by Johannes Müller Argoviensis Taxa described in 1891 {{Parmeliaceae-stub ...
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Synonym (taxonomy)
In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The Botanical nomenclature, botanical and Zoological nomenclature, zoological codes of nomenclature treat the concept of synonymy differently. * In nomenclature, botanical nomenclature, a synonym is a Binomial nomenclature, scientific name that applies to a taxon that now goes by a different scientific name. For example, Carl Linnaeus, Linnaeus was the first to give a scientific name (under the currently used system of scientific nomenclature) to the Norway spruce, which he called ''Pinus abies''. This name is no longer in use, so it is now a synonym of the current scientific name, ''Picea abies''. * In zoology, moving a species from one genus to another results in a different Binomial nomenclature, binomen, but the name is considered an alternative combination rather than a synonym. The concept of synonymy in zoology is reserved f ...
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Evolutionary Divergence
Divergent evolution or divergent selection is the accumulation of differences between closely related populations within a species, sometimes leading to speciation. Divergent evolution is typically exhibited when two populations become separated by a geographic barrier (such as in allopatric or peripatric speciation) and experience different selective pressures that cause adaptations. After many generations and continual evolution, the populations become less able to interbreed with one another. The American naturalist J. T. Gulick (1832–1923) was the first to use the term "divergent evolution", with its use becoming widespread in modern evolutionary literature. Examples of divergence in nature are the adaptive radiation of the finches of the Galápagos, changes in mobbing behavior of the kittiwake, and the evolution of the modern-day dog from the wolf. The term can also be applied in molecular evolution, such as to proteins that derive from homologous genes. Both ortho ...
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Molecular Phylogeny
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to determine the processes by which diversity among species has been achieved. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree. Molecular phylogenetics is one aspect of molecular systematics, a broader term that also includes the use of molecular data in taxonomy and biogeography. Molecular phylogenetics and molecular evolution correlate. Molecular evolution is the process of selective changes (mutations) at a molecular level (genes, proteins, etc.) throughout various branches in the tree of life (evolution). Molecular phylogenetics makes inferences of the evolutionary relationships that arise due to molecular evolution and results in the construction of a phylogenetic tree. History The theoretical framew ...
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