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Tropeognathus Mesembrinus
''Tropeognathus'' (meaning "keel jaw") is a genus of large pterosaurs from the late Early Cretaceous of South America. This genus is considered to be a member of the family Anhangueridae, however, several studies have also recovered it within another family called Ornithocheiridae. Both of these families are diverse groups of pterosaurs known for their keel-tipped snouts and large size. ''Tropeognathus'' is regarded as the largest pterosaur found in the Southern Hemisphere, only rivaled by the huge azhdarchids. The type and only species is ''Tropeognathus mesembrinus''. Fossil remains of ''Tropeognathus'' have been recovered from the Romualdo Formation, which is a Lagerstätte located in the Santana Group of the Araripe Basin in northeastern Brazil. Discovery and naming In the 1980s the German paleontology museum '' Bayerische Staatssammlung für Paläontologie und historische Geologie'' in Munich acquired a pterosaur skull from Brazilian fossil dealers that had probabl ...
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Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 Megaannum#SI prefix multipliers, Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian–Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining Anoxic event#Cretaceous, early Aptian Oceanic Anoxic Event 1a (OAE1a) was dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java Plateau, Ontong Java-Manihiki Plateau, Manihiki-Hikurangi Plateau, Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Onto ...
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Lagerstätte
A Fossil-Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that preserves an exceptionally high amount of palaeontological information. ''Konzentrat-Lagerstätten'' preserve a high concentration of fossils, while ''Konservat-Lagerstätten'' offer exceptional fossil preservation, sometimes including preserved soft tissues. ''Konservat-Lagerstätten'' may have resulted from carcass burial in an anoxic environment with minimal bacteria, thus delaying the decomposition of both gross and fine biological features until long after a durable impression was created in the surrounding matrix. ''Fossil-Lagerstätten'' spans geological time from the Neoproterozoic era to the present. Worldwide, some of the best examples of near-perfect fossilization are the Cambrian Maotianshan shales and Burgess Shale, the Ordovician Soom Shale, the Silurian Waukesha Biota, the Devonian Hunsrück Slates and Gogo Formation, the Ca ...
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Holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany and mycology, an isotype is a duplicate of the holotype, generally pieces from the same individual plant or samples from the same genetic individual. A holotype is not necessarily "ty ...
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Koine
Koine Greek (, ), also variously known as Hellenistic Greek, common Attic, the Alexandrian dialect, Biblical Greek, Septuagint Greek or New Testament Greek, was the common supra-regional form of Greek spoken and written during the Hellenistic period, the Roman Empire and the early Byzantine Empire. It evolved from the spread of Greek following the conquests of Alexander the Great in the fourth century BC, and served as the lingua franca of much of the Mediterranean region and the Middle East during the following centuries . It was based mainly on Attic and related Ionic speech forms, with various admixtures brought about through dialect levelling with other varieties. Koine Greek included styles ranging from conservative literary forms to the spoken vernaculars of the time. As the dominant language of the Byzantine Empire, it developed further into Medieval Greek, which then turned into Modern Greek. Literary Koine was the medium of much post-classical Greek literary and scho ...
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Specific Name (zoology)
In zoological nomenclature, the specific name (also specific epithet, species epithet, or epitheton) is the second part (the second name) within the scientific name of a species (a binomen). The first part of the name of a species is the name of the genus or the generic name. The rules and regulations governing the giving of a new species name are explained in the article species description. For example, the scientific name for humans is ''Homo sapiens'', which is the species name, consisting of two names: ''Homo'' is the " generic name" (the name of the genus) and ''sapiens'' is the "specific name". Etymology Historically, ''specific name'' referred to the combination of what are now called the generic and specific names. Carl Linnaeus, who formalized binomial nomenclature, made explicit distinctions between specific, generic, and trivial names. The generic name was that of the genus, the first in the binomial, the trivial name was the second name in the binomial, and the ...
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Type Species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological Type (biology), type wiktionary:en:specimen, specimen (or specimens). Article 67.1 A similar concept is used for suprageneric groups and called a type genus. In botanical nomenclature, these terms have no formal standing under the International Code of Nomenclature for algae, fungi, and plants, code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen (or, rarely, an illustration) which is also the type of a species name. The species name with that type can also be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have suc ...
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Chapada Do Araripe
The Chapada do Araripe, also known as the Serra do Araripe, is a ''chapada'' (plateau) in northeastern Brazil. The chapada forms the boundary of Ceará and Pernambuco states, and forms the watershed between the Jaguaribe River of Ceará, which flows northward into the Atlantic, and the much larger basin of the São Francisco River of Pernambuco and neighboring states, which drains eastward into the Atlantic. The Santana Formation, which is rich with fossils from the early Cretaceous, lies at the base of the chapada. The Araripe manakin (''Antilophia bokermanni''), a bird only described in 1998, is endemic to Chapada do Araripe. Etymology The word "Araripe" stems from the Tupi word ''ararype'', meaning "macaw Macaws are a group of Neotropical parrot, New World parrots that are long-tailed and often colorful, in the Tribe (biology), tribe Arini (tribe), Arini. They are popular in aviculture or as companion parrots, although there are conservation con ... at the river" (''ar ...
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Ceará
Ceará (, ) is one of the 26 states of Brazil, located in the Northeast Region, Brazil, northeastern part of the country, on the Atlantic Ocean, Atlantic coast. It is the List of Brazilian states by population, eighth-largest Brazilian State by population and the List of Brazilian states by area, 17th by area. It is also one of the main tourist destinations in Brazil. The List of capitals in Brazil, state capital is the city of Fortaleza, the country's fourth most populous city. The state has 4.3% of the Brazilian population and produces 2.1% of the Brazilian GDP. It is divided into 184 municipalities. Literally, the name ''Ceará'' means "sings the Aratinga, jandaia". According to José de Alencar, one of the most important writers of Brazil and an authority in Tupi Guaraní, ''Ceará'' means turquoise or green waters. The state is best known for its extensive coastline, with of sand. There are also mountains and valleys producing tropical fruits. To the south, on the border o ...
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Munich
Munich is the capital and most populous city of Bavaria, Germany. As of 30 November 2024, its population was 1,604,384, making it the third-largest city in Germany after Berlin and Hamburg. Munich is the largest city in Germany that is not a state of its own. It ranks as the 11th-largest city in the European Union. The metropolitan area has around 3 million inhabitants, and the broader Munich Metropolitan Region is home to about 6.2 million people. It is the List of EU metropolitan regions by GDP#2021 ranking of top four German metropolitan regions, third largest metropolitan region by GDP in the European Union. Munich is located on the river Isar north of the Alps. It is the seat of the Upper Bavaria, Upper Bavarian administrative region. With 4,500 people per km2, Munich is Germany's most densely populated municipality. It is also the second-largest city in the Bavarian language, Bavarian dialect area after Vienna. The first record of Munich dates to 1158. The city ha ...
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Paleontology
Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geologic time, and assess the interactions between prehistoric organisms and their natural environment. While paleontological observations are known from at least the 6th century BC, the foundation of paleontology as a science dates back to the work of Georges Cuvier in 1796. Cuvier demonstrated evidence for the concept of extinction and how life of the past was not necessarily the same as that of the present. The field developed rapidly over the course of the following decades, and the French word ''paléontologie'' was introduced for the study in 1822, which was derived from the Ancient Greek word for "ancient" and words describing relatedness and a field of study. Further advances in the field accompanied the work of Charles Darwin ...
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Tropeognathus
''Tropeognathus'' (meaning "keel jaw") is a genus of large pterosaurs from the late Early Cretaceous of South America. This genus is considered to be a member of the family Anhangueridae, however, several studies have also recovered it within another family called Ornithocheiridae. Both of these families are diverse groups of pterosaurs known for their keel-tipped snouts and large size. ''Tropeognathus'' is regarded as the largest pterosaur found in the Southern Hemisphere, only rivaled by the huge azhdarchids. The type and only species is ''Tropeognathus mesembrinus''. Fossil remains of ''Tropeognathus'' have been recovered from the Romualdo Formation, which is a Lagerstätte located in the Santana Group of the Araripe Basin in northeastern Brazil. Discovery and naming In the 1980s the German paleontology museum ''Bayerische Staatssammlung für Paläontologie und historische Geologie'' in Munich acquired a pterosaur skull from Brazilian fossil dealers that had probably been ...
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