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In
zoological nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its formal author, t ...
, a type species (''species typica'') is the
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
name with which the name of a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
or
subgenus In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
is considered to be permanently taxonomically associated, i.e., the species that contains the biological
type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * ...
specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository f ...
(or specimens). Article 67.1 A similar concept is used for suprageneric groups and called a
type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...
. In
botanical nomenclature Botanical nomenclature is the formal, scientific naming of plants. It is related to, but distinct from taxonomy (biology), taxonomy. Plant taxonomy is concerned with grouping and classifying plants; Botany, botanical nomenclature then provides na ...
, these terms have no formal standing under the code of nomenclature, but are sometimes borrowed from zoological nomenclature. In botany, the type of a genus name is a specimen (or, rarely, an illustration) which is also the type of a species name. The species name with that type can also be referred to as the type of the genus name. Names of genus and family ranks, the various subdivisions of those ranks, and some higher-rank names based on genus names, have such types.''International Code of Nomenclature for algae, fungi, and plants'' (ICN)
articles 7 through 10 (Melbourne Code, 2012)
In
bacteriology Bacteriology is the branch and specialty of biology that studies the Morphology (biology), morphology, ecology, genetics and biochemistry of bacteria as well as many other aspects related to them. This subdivision of microbiology involves the iden ...
, a type species is assigned for each genus. Whether or not currently recognized as valid, every named genus or subgenus in zoology is theoretically associated with a type species. In practice, however, there is a backlog of untypified names defined in older publications when it was not required to specify a type.


Use in zoology

A type species is both a concept and a practical system that is used in the
classification Classification is the activity of assigning objects to some pre-existing classes or categories. This is distinct from the task of establishing the classes themselves (for example through cluster analysis). Examples include diagnostic tests, identif ...
and nomenclature (naming) of animals. The "type species" represents the reference species and thus "definition" for a particular genus name. Whenever a taxon containing multiple species must be divided into more than one genus, the type species automatically assigns the name of the original taxon to one of the resulting new taxa, the one that includes the type species. The term "type species" is regulated in
zoological nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its formal author, t ...
by article 42.3 of the ''
International Code of Zoological Nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted Convention (norm), convention in zoology that rules the formal scientific name, scientific naming of organisms treated as animals. It is also informally known as the I ...
'', which defines a type species as the
name-bearing type Under the ''International Code of Zoological Nomenclature'' (''Code''), the name-bearing type or onomatophore is the biological type that determines the application of a name. Each animal taxon regulated by the ''Code'' at least potentially has a na ...
of the name of a genus or subgenus (a " genus-group name"). In the Glossary, type species is defined as The type species permanently attaches a formal name (the generic name) to a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
by providing just one
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
within that genus to which the genus name is permanently linked (i.e. the genus must include that species if it is to bear the name). The species name in turn is fixed, in theory, to a type specimen. For example, the type species for the
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial molluscs, terrestrial gastropod mollusks that have gastropod shell, shel ...
genus '' Monacha'' is ''Helix cartusiana'', the name under which the species was first described, known as ''
Monacha cartusiana ''Monacha cartusiana'' is a species of small air-breathing land snail, a terrestrial molluscs, terrestrial Eupulmonata, eupulmonate gastropod mollusc in the family Hygromiidae, the hairy snails and their allies. This is the type species of the ...
'' when placed in the genus ''Monacha''. That genus is currently placed within the family
Hygromiidae Hygromiidae is a taxonomic family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea.Helicellidae Ihering, 1909 Anatomy Some snails in genera within this family create and ...
. The type genus for that family is the genus ''
Hygromia ''Hygromia'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Hygromiidae, the hairy snails and their allies.MolluscaBase eds. (2021). MolluscaBase. Hygromia Risso, 1826. Accessed through: World Re ...
''. The concept of the type species in zoology was introduced by
Pierre André Latreille Pierre André Latreille (; 29 November 1762 – 6 February 1833) was a French zoology, zoologist, specialising in arthropods. Having trained as a Roman Catholic priest before the French Revolution, Latreille was imprisoned, and only regained hi ...
.


Citing

The ''International Code of Zoological Nomenclature'' states that the original name (binomen) of the type species should always be cited. It gives an example in Article 67.1. ''Astacus marinus'' was later designated as the type species of the genus ''Homarus'', thus giving it the name '' Homarus marinus'' . However, the type species of ''Homarus'' should always be cited using its original name, i.e. ''Astacus marinus'' , even though that is a
junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...
of ''Cancer grammarius'' . Although the ''
International Code of Nomenclature for algae, fungi, and plants The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN or ICNafp) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all tho ...
'' does not contain the same explicit statement, examples make it clear that the original name is used, so that the "type species" of a genus name need not have a name within that genus. Thus in Article 10, Ex. 3, the type of the genus name ''Elodes'' is quoted as the type of the species name '' Hypericum aegypticum'', not as the type of the species name ''Elodes aegyptica''. (''Elodes'' is not now considered distinct from ''
Hypericum ''Hypericum'' is a genus of flowering plants in the family (biology), family Hypericaceae (formerly considered a subfamily of Clusiaceae). The genus has a nearly worldwide distribution, missing only from tropical lowlands, deserts and polar re ...
''.)


See also

*
Glossary of scientific naming This is a list of terms and symbols used in scientific names for organisms, and in describing the names. For proper parts of the names themselves, see List of Latin and Greek words commonly used in systematic names. Many of the abbreviations are ...
* Genetypes – genetic sequence data from type specimens. *
Holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
*
Paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype (biology), isotype ...
* Principle of typification *
Type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * ...
(and
type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...
) *
Type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...


References

{{Authority control Botanical nomenclature Taxonomy (biology) Zoological nomenclature