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Trigonotarbida
The order Trigonotarbida is a group of extinct arachnids whose fossil record extends from the late Silurian to the early Permian ( Pridoli to Sakmarian).Dunlop, J. A., Penney, D. & Jekel, D. 2020A summary list of fossil spiders and their relatives In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch , version 20.5 These animals are known from several localities in Europe and North America, as well as a single record from Argentina. Trigonotarbids can be envisaged as spider-like arachnids, but without silk-producing spinnerets. They ranged in size from a few millimetres to a few centimetres in body length and had segmented abdomens (opisthosoma), with the dorsal exoskeleton ( tergites) across the backs of the animals' abdomens, which were characteristically divided into three or five separate plates. Probably living as predators on other arthropods, some later trigonotarbid species were quite heavily armoured and protected themselves with spines and tub ...
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Arachnid
Arachnids are arthropods in the Class (biology), class Arachnida () of the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, opiliones, harvestmen, Solifugae, camel spiders, Amblypygi, whip spiders and Uropygi, vinegaroons. Adult arachnids have eight Arthropod leg, legs attached to the cephalothorax. In some species the frontmost pair of legs has converted to a sensory function, while in others, different appendages can grow large enough to take on the appearance of extra pairs of legs. Almost all Extant taxon, extant arachnids are terrestrial animal, terrestrial, living mainly on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well. They comprise over 110,000 named species, of which 51,000 are species of spiders. The term is derived from the Ancient Greek, Greek word (''aráchnē'', 'spider'), from the myth of the hubristic human weaver Arachne, w ...
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Anthracomartidae
Anthracomartidae, first described by Haase, 1890, is a family of the extinct arachnid order Trigonotarbida. It is likely to be most closely related to the Archaeomartidae, based on a 2014 cladistic analysis, within the clade (Palaeocharinus (Archaeomartidae + Anthracomartidae)). Many specimens come from siderite nodules of late Carboniferous The Carboniferous ( ) is a Geologic time scale, geologic period and System (stratigraphy), system of the Paleozoic era (geology), era that spans 60 million years, from the end of the Devonian Period Ma (million years ago) to the beginning of the ... age in Europe and North America. Genera Anthracomartidae contains the genera : *'' Anthracomartus'' Karsch, 1882 *'' Brachypyge'' Woodward, 1878 *'' Maiocercus'' Pocock, 1911 * The genus also was previously considered to contain other genera before they were later synonymized with Anthracomartus and various species within. These include: * '' Brachylycosa'' Frič, 1904 * '' Cleptomartus ...
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Maiocercus
''Maiocercus celticus'' is a species of early trigonotarbid arachnid from the Upper Carboniferous of Westhoughton, Lancashire, UK. The species was first described in 1902, with a "new species" being described in 1911 (''M. orbicularis'') which has been proven as being a junior synonym of ''M. celticus''. ''M. celticus'' is the type species of the genus ''Maiocercus''. Originally zoologist Reginald Innes Pocock compared ''M. celticus'' to Brachypyge, with later evidence showing that ''Brachypyge'' had "opisthosoma which were much longer than wide; with the pleural laminæ of the second and third pleura-bearing terga being inclined slightly backwards" (''Brachypyge'') with ''Maiocercus'' having the "opisthosoma much wider than long; the pleural laminæ of the first, second, third, and fourth sterna being inclined slightly forwards". The original drawing which showed ''Maiocercus'' described a pitting on the underside of the slightly forwarded laminæ, with a non-uniform concavi ...
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Carboniferous
The Carboniferous ( ) is a Geologic time scale, geologic period and System (stratigraphy), system of the Paleozoic era (geology), era that spans 60 million years, from the end of the Devonian Period Ma (million years ago) to the beginning of the Permian Period, Ma. It is the fifth and penultimate period of the Paleozoic era and the fifth period of the Phanerozoic eon (geology), eon. In North America, the Carboniferous is often treated as two separate geological periods, the earlier Mississippian (geology), Mississippian and the later Pennsylvanian (geology), Pennsylvanian. The name ''Carboniferous'' means "coal-bearing", from the Latin ("coal") and ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern "system" names, it was coined by geologists William Conybeare (geologist), William Conybeare and William Phillips (geologist), William Phillips in 1822, based on a study of the British rock succession. Carboniferous is the per ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods, arachnids and hexapods) ...
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Eophrynidae
Eophrynidae is a family of the extinct arachnid order Trigonotarbida. Eophrynids lived during the Carboniferous The Carboniferous ( ) is a Geologic time scale, geologic period and System (stratigraphy), system of the Paleozoic era (geology), era that spans 60 million years, from the end of the Devonian Period Ma (million years ago) to the beginning of the ... period in what is now modern Europe and North America. The family is probably found within the "eophrynid assemblage" clade: (''Aphantomartus'' (''Alkenia'' (''Pseudokreischeria'' (''Kreischeria'' (''Eophrynus'' + ''Pleophrynus''))))). Genera *'' Areomartus'' Petrunkevitch, 1913 *'' Eophrynus'' Woodward, 1871 *'' Nyranytarbus'' Harvey & Selden, 1995 *'' Petrovicia'' Frič, 1904 *'' Planomartus'' Petrunkevitch, 1953 *'' Pleophrynus'' Petrunkevitch, 1945 *'' Pocononia'' Petrunkevitch, 1953 *'' Somaspidion'' Jux, 1982 *'' Stenotrogulus'' Frič, 1904 *'' Vratislavia'' Frič, 1904 References Trigonotarbid ...
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Late Silurian
The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became ...
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Palaeocharinus
''Palaeocharinus'' is a genus of extinct trigonotarbid arachnids known from the Devonian of western Europe. The genus was first found and described in the Rhynie chert in the 1920s by Arthur Stanley Hirst and S. Maulik. The family to which the genus belongs may be paraphyletic. Species * ''Palaeocharinus calmani'' (Hirst, 1923) – Early Devonian The Early Devonian is the first of three Epoch (geology), epochs comprising the Devonian period, corresponding to the Lower Devonian Series (stratigraphy), series. It lasted from and began with the Lochkovian Stage , which was followed by the Pr ..., Scotland * ''Palaeocharinus hornei'' (Hirst, 1923) – Early Devonian, Scotland * ''Palaeocharinus kidstoni'' (Hirst, 1923) – Early Devonian, Scotland * ''Palaeocharinus rhyniensis'' (Hirst, 1923) – Early Devonian, Scotland * ''Palaeocharinus scourfieldi'' (Hirst, 1923) – Early Devonian, Scotland * ''Palaeocharinus tuberculatus'' (Fayers, Dunlop & Trewin, 2005) – Early Devonia ...
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Eophrynus
''Eophrynus'' is an extinct genus of arachnids from the extinct order Trigonotarbida, which lived during the Late Carboniferous period in Europe. The genus was first described in 1871 by Henry Woodward (geologist). The name comes from ''Eo'', meaning 'dawn', and '' Phrynus'', an extant genus of whip spider (order Amblypygi). Two species have been recognised: * '' Eophrynus prestvicii'' in England * '' Eophrynus udus'' in Germany Species of ''Eophrynus'', as with other tribonotarbids, were similar to modern spiders but could not produce silk and the back-half of their body was made up of small plates. The English species, ''E. prestvici'', is known from a handful of good quality fossils preserved inside siderite concretions. Recent X-ray An X-ray (also known in many languages as Röntgen radiation) is a form of high-energy electromagnetic radiation with a wavelength shorter than those of ultraviolet rays and longer than those of gamma rays. Roughly, X-rays have a wavelengt ...
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Sakmarian
In the geologic timescale, the Sakmarian is an age or stage of the Permian period. It is a subdivision of the Cisuralian Epoch or Series. The Sakmarian lasted between 293.52 and million years ago (Ma). It was preceded by the Asselian and followed by the Artinskian.; 2004: ''A Geologic Time Scale 2004'', Cambridge University Press Stratigraphy The Sakmarian Stage is named after the Sakmara River in the Ural Mountains, a tributary to the Ural River. The stage was introduced into scientific literature by Alexander Karpinsky in 1874. In Russian stratigraphy, it originally formed a substage of the Artinskian Stage. Currently, the ICS (International Commission on Stratigraphy) uses it as an independent stage in its international geologic timescale. The base of the Sakmarian Stage is defined by the first appearance of conodont species '' Streptognathodus postfusus'' in the fossil record. A global reference profile for the stage's base (a GSSP A Global Boundary Stratotype ...
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Arthropod
Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metamerism (biology), metameric) Segmentation (biology), segments, and paired jointed appendages. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. They form an extremely diverse group of up to ten million species. Haemolymph is the analogue of blood for most arthropods. An arthropod has an open circulatory system, with a body cavity called a haemocoel through which haemolymph circulates to the interior Organ (anatomy), organs. Like their exteriors, the internal organs of arthropods are generally built of repeated segments. They have ladder-like nervous systems, with paired Anatomical terms of location#Dorsal and ventral, ventral Ventral nerve cord, nerve cord ...
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